7.16. THOMISIDAE (INCL. PHILODROMIDAE) 155
156 CHAPTER 7. KEYS AND DESCRIPTIONS separated between these bands and has the form of a weakly curved lamella (Fig. 183).
The male palpus bears two apophyses (outer and ventral); the ventral apophysis is not broad and long and is more or less formed like an inclined cut-out lamella. The dorsal apophysis is markedly thick, long, tapering towards the end and directed forward (Fig. 182). Carapace length 22.5 mm.
Habitat: On vegetables, mostly on cabbage with corresponding color (green or yellowish white); on grass. They feed on pest insects of cabbage and gardens: aphids, bugs, larvae of pierids and others.
Distribution: Holarctic (except the far north). In Georgia: Kutaisi, Zestaponi (= Qvirila). (Kulczy«ski 1895), Tbilisi, Gori, Khashuri, Borjomi, Telavi, Sighnaghi, Lagodekhi (Mt. Ninigori), Satshkhere, Kharagauli, Bagh-dati, Shovi, Poti (Mcheidze 19411963).
Taxonomy: Platnick (2013): Philodromus aureolus (Clerck, 1757). Mchei-dze (1997) also lists the synonym Ph. canadensis Emerton, 1917.
7. Philodromus aureolus caespiticola (Walckenaer, 1805)
Kulczy«ski (1895) found this species in Gelati. We could not nd it in our material.
Taxonomy: Platnick (2013): Philodromus cespitum (Walckenaer, 1802)56. 7.16.2 Thanatus C. L. Koch, 1837
Key to species
1(6) Males. . . 2 2(3) Carapace broader than abdomen, almost rounded. Dark brown part of the carapace with characteristic markings; its sides with broad, almost semilunar, dark brown markings. Median line of the eye region with three short brown bands but in the central part with a large mark of irregular shape. Abdomen elongated oval, light yellow, at the median line with light brown, dark brown and red markings (Fig. 186). . . 1. Th. lineatipes (p. 157) 3(2) Coloration of carapace and abdomen not like this. . . 4 4(5) Palpus tibia apophysis near the tip broadened (Fig. 193). Cara-pace brown with two longitudinal light bands and light lateral edge. Abdomen pink-yellow with a black lanceolate mark in its anterior part and two longitudinal brown bands or with posterior sharp raspberry-colored bands. Female body length 6.57.5 mm, male 6 mm. . . 2. Th. arenarius (p. 157)
56In Mcheidze (1997) sic 'Philodzomus aureolus (Cl.) Sub sp. caespiticola (Walck., 1805)', lapsi.
7.16. THOMISIDAE (INCL. PHILODROMIDAE) 157 5(4) Palpus tibia apophysis near the tip gradually tapering (Fig. 191).
Carapace light brown with two longitudinal dark bands. Ab-domen pinkish-yellow, sometimes distinctly pink with a lanceo-late black or a brown band in the posterior half. Male coloration darker than in female. Female body length 6.58 mm, male 5.5 mm. . . 3. Th. formicinus (p. 159) 6(1) Females. . . 7 7(8) Epigynal median plate narrow, not broader in its posterior half (Fig. 190). Body length 58 mm. . 3. Th. formicinus (p. 159) 8(7) Epigynal plate dierent. . . 9 9(10) Epigynal plate laterally bordered by a narrow chitinized arch (Fig.
192). Body length 79 mm. . . 2. Th. arenarius (p. 157) 10(9) Epigynal plate not bordered by such an arch. . . 11 11(12) Median plate of the epigyne broad and somewhat narrowed in its posterior half (Fig. 187). . . 1. Th. lineatipes (p. 157) 12(11) Median plate of the epigyne narrow and in its anterior part cone-like narrowed. The form of the epigynal pits is similar to the shape of wrongly paired shoes (right-left and left-right) (Fig.
188). . . 4. Th. imbecilus (p. 159) 1. Thanatus lineatipes Simon, 1870
Description: The sternum of the female is blackish. The clypeus is narrow and brighter than the legs, which are long, with lineation and, especially on the femur, with two brown bands. The epigynal plate is semi-circular (Fig.
187). The male bulbus bears a cut-o apophysis, which bears a small conical bulging part.
Habitat: In short grass in rocky places.
Distribution: Spain, Portugal, Tunis, Syria. In Georgia: Tbilisi (Mchei-dze 1959). First record in the former USSR.
Taxonomy: Platnick (2013): Thanatus lineatipes Simon, 187057. 2. Thanatus arenarius L. Koch, 1872
Description: Female body length 6.57.5 mm, male 6 mm.
„ The carapace is brownish red, sometimes carmine red and brighter than in the male; with two elongated bands, which are widened near the tip.
The ME are uniform, the AE form a weakly curved line.
The anterior part of the longish-oval abdomen is hardly distinguishable brown red and covered with light brown setae as well as with a median black band in the anterior half, which is rectangular in its anterior part and tapering in the posterior part. This band is covered with brownish red setae and is often bordered by a narrow white band. In the second half this band is
57In Mcheidze (1997) sic: 'Thanatus liniatipes (E. Simon, 1870)', lapsi.
158 CHAPTER 7. KEYS AND DESCRIPTIONS
Fig. 179185. Thomisidae (Philodromidae), Philodromus. Ph.
histrio: 179 ventral male palp; 179 a lateral male palp; 180 habi-tus; 180 a epigyne. Ph. dispar: 181 male palp. Ph. aureolus: 182 palpal apophysis; 183 epigyne. Ph. rufus: 184 palpal apophysis; 184 a epigyne. Ph. margaritatus: 185 epigyne.
weakly curved and ends above the spinnerets. The extremities are brownish red and covered with whitish setae. The epigyne forms a large brown-red depression (Fig. 192).
ƒ The carapace is darkened brownish-red with two longitudinal light lines, directed from the posterior part to the sides of the eye region. Its posterior part is weakly narrowed and covered with light yellow setae. The cephalic region is very short and blunt, its surface almost at, the thoracic part is not raised but with well visible bands on the sides, which do, however, not extend all along the rst half of the carapace. The PME are smaller and more condensed than the LE. The legs are mostly bright brown-red. Palpus brownish red (Fig. 193).
Habitat: In high grass, shrubs, rarely under rocks. Steppe species.
Distribution: Europe, Turkey, in the former USSR in Kharatov region, Voronezh region, Ukraine, Urals and Central Asia. In Georgia: Tbilisi (Simon 1899), Kiketi, Betania, Poti (Mcheidze 1968).
Taxonomy: Platnick (2013): Thanatus arenarius L. Koch, 187258. She also lists the synonym T. mundus Pickard-Cambridge, 1873.
58In Mcheidze (1997) sic: 'Thor., 1872', lapsus.
7.16. THOMISIDAE (INCL. PHILODROMIDAE) 159 3. Thanatus formicinus (Clerck, 1757)
Description: Female body length 6.58 mm, males 5.5 mm. The carapace is light brown, with two long dark bands. Sternum and abdomen are pinkish-yellow; the abdomen is sometimes shining pink, with a black lanceolate mark in the anterior half and two long, pink or brown bands in the posterior half.
The coloration is darker in males compared to females. Epigyne like Fig.
190. Male palpus like Fig. 191.
Habitat: On the ground, in grass.
Distribution: Central and southern European countries, North America, in the former USSR everywhere except the extreme North. In Georgia:
Mestia, Bakhmaro, Adigeni (Mcheidze 1940). First record in the Transcau-casus.
Taxonomy: Platnick (2013): Thanatus formicinus (Clerck, 1757).
4. Thanatus imbecillus L. Koch, 1878
Description: „ The sides of the brown-yellow carapace are black and mar-morated; a long band, which is enlarged in its upper part and whitish-yellow towards the AER orginates from the posterior carapace slope. The cephalic region bears two reddish-brown bands, which unite in the posterior part and show a line of the same color between them.
The palpus is bulged, light brown; its center is reddish brown. Most of the setae are white and wing-shaped; the ventral and lateral parts of the cephalic region bear a trail of yellow setae. Both eye rows are curved forward, especially the posterior row. The sternum is light brown and covered with white setae as well as black sprinkles. Metatarsus and tarsus I bear a scopula.
The legs are thin, long and covered with wing-shaped setae.
The abdomen is elongated, rounded, its posterior part is pointed. The main color of the abdomen is claycolored-brownish-yellow, with a black dor-sal mark. The white fringe is narrow, with a rhomboid mark originating from the basis and not reaching the central part (Fig. 189). The venter bears four elongated brown lines, of which the two central lines unite in the posterior part, terminating in a point. The spinnerets are brown-yellow white. Epigyne like Fig. 188.
Habitat: In grass.
Distribution: Turkey. In the former USSR in the Caspian Region, Tran-scaucasus. In Georgia: Tbilisi (Kulczy«ski 1899), Tbilisi, Akhalkalaki, Gori (Mcheidze 1970).
Taxonomy: Platnick (2013): Thanatus imbecillus L. Koch, 187859.
59In Mcheidze (1997) sic: 'imbecilus', lapsus, and author with brackets.
160 CHAPTER 7. KEYS AND DESCRIPTIONS
Figs. 186193. Thomisidae (Philodromidae), Thanatus. T. lineatipes:
186 habitus; 187 epigyne. T. imbecilus: 188 epigyne; 189 habitus. T.
formicinus: 190 epigyne; 191 male palpus. T. arenarius: 192 epigyne;
193 male palpus.
7.16. THOMISIDAE (INCL. PHILODROMIDAE) 161