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Lukas und Marike. Mein herzlicher Dank richtet sich an Steffen Höll für seinen großen Einsatz bei der Gewinnung von Sequenzdaten und vor allem für sein besonderes Interesse an der vorliegenden Thematik und die daraus resultierenden gemeinsamen Diskussionen.

Bei der phylogenetischen Auswertung der molekularen Datensätze haben mich außerdem Anna Hundsdörfer (Senckenberg Museum für Tierkunde Dresden) und Lars Opgenoorth (Universität Marburg) unterstützt. Ich hoffe, dass ich die Möglichkeit haben werde, mit diesem hervorragenden Team noch viele weitere Laufkäfer-Phylogeographien zu erarbeiten.

Die Zeit, die ich mit Rucksack und Zelt im Himalaya fernab der Zivilisation verbracht habe, summiert sich auf Jahre. Dass alle diese Expeditionen mit wertvollen Materialsammlungen endeten und trotz aller erdenklichen Abenteuer immer gut ausgingen, habe ich nicht zuletzt meinen nepalesischen Helfern zu verdanken, allen voran Santos Tamang. Auch mein langjähriger Reisegefährte Olaf Jäger (Museum für Tierkunde Dresden) hat wesentlich zum Gelingen dieser Expeditionen beigetragen. Der Anstoß für eine Erforschung der Laufkäfer Nepals geht auf meinen Freund Matthias Hartmann (Naturkundemuseum Erfurt) zurück. Von Matthias und Olaf erhielt ich alle nur denkbare Unterstützung, die ich über die vielen Jahre Freizeitforschung im Himalaya benötigte. In Tibet haben mir neben Torstein vor allem Pubu, Sonam Tso und Tsering geholfen, meine Feldarbeiten erfolgreich durchzuführen.

Last but not least, bedanke ich mich bei den Mitarbeitern im Fachbereich Geographie der Universität Marburg für die unkomplizierte Gewährung der vielen Extrawünsche aufgrund meines Arbeitsplatzes im fernen Rostock. Ganz besonders herzlich bedanke ich mich bei Frau Christina Philippi für ihre beständige Unterstützung und unproblematische Hilfe zur Bewältigung der Bürokratie. Frau Christiane Enderle hat außerdem für die vorliegende Studie hervorragende Kartengrundlagen erarbeitet. Auf diesen basieren die Abbildungen 8-9, 13-14 und 21-28.

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Publikation I

Schmidt, J.

Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau

(Coleoptera: Carabidae: Trechini).

Zootaxa (2009) 2178: 1-72.

Accepted by P. Johnson: 2 Jul. 2009; published: 6 Aug. 2009

ISSN1175-5334(online edition) Copyright © 2009 · Magnolia Press

ZOOTAXA

Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern

central Tibetan Plateau (Coleoptera: Carabidae: Trechini)

JOACHIM SCHMIDT

Faculty of Geography, University of Marburg, Deutschhausstrasse 10, 35037 Marburg, Germany.

Email: schmidt@agonum.de

Magnolia Press Auckland, New Zealand

2178

SCHMIDT

2 · Zootaxa 2178 © 2009 Magnolia Press

laya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) (Zootaxa 2178)

72 pp.; 30 cm.

6 Aug. 2009

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Trechus from southern central Tibet

Abstract ... 4 Introduction ... 5 Material and methods ... 5 Taxonomy ... 7 A Key to Trechus species of the Tibetan Himalaya and the Southern Tibetan Plateau ... 7 The Trechus thibetanus group ... 11 Trechus eutrechoides Deuve, 1992 ... 13 Trechus eutrechoides mondaensis Deuve, 1997... 13 Trechus thorongiensis Schmidt, 1994 ... 17 Trechus thibetanus Jeannel, 1928... 17 Trechus namtsoensis sp. n... 18 Trechus dongulaensis sp. n. ... 19 Trechus glabratus sp. n. ... 20 The Trechus wrzecionkoi group ... 21 Trechus wrzecionkoi Deuve, 1996 ... 22 Trechus korae sp. n. ... 22 Trechus martinae sp. n. ... 23 The Trechus franzianus group ... 24 Trechus pumoensis Deuve, 1997 ... 25 Trechus tilitshoensis Schmidt, 1994 ... 25 Trechus franzianus Mateu & Deuve, 1979... 26 Trechus muguensis sp. n. ... 27 Trechus eremita sp. n. ... 28 Trechus aedeagalis sp. n. ... 29 Trechus sculptipennis sp. n. ... 35 The Trechus dacatraianus group ... 37 Trechus damchungensis Deuve, 1997 ... 37 Trechus hodeberti Deuve, 1997 ... 38 Trechus bastropi sp. n. ... 38 Trechus mieheorum sp. n. ... 40 The Trechus solhoeyi group ... 41 Trechus solhoeyi sp. n. ... 41 The Trechus antonini group ... 42 Trechus budhaensis sp. n. ... 43 Trechus yeti sp. n. ... 44 Trechus antonini Deuve, 1997 ... 45 Trechus religiosus sp. n. ... 46 Trechus yak sp. n. ... 47 Trechus yak shogulaensis ssp. n. ... 48 Trechus folwarcznyi Deuve, 1997 ... 49 Trechus tsampa sp. n. ... 49 Trechus rarus sp. n. ... 50 Trechus singularis sp. n. ... 51 Trechus tseringi sp. n. ... 52 Trechus astrophilus sp. n. ... 53 Trechus lama sp. n. ... 55 The Trechus chaklaensis group ... 56 Trechus chaklaensis sp. n. ... 57 The Trechus stratiotes group ... 58 Trechus stratiotes sp. n. ... 58 Trechus stratiotes malikasthana ssp. n. ... 60 The Trechus rolwalingensis group ... 61