Phylogeny and taxonomic revision of the genus Eristalopsis Evenhuis (Diptera: Bombyliidae: Bombyliinae)
XUANKUN LI *, 1, 2 & DAVID K. YEATES1
1 Australian National Insect Collection, CSIRO National Research Collections Australia, Canberra, ACT 2601, Australia; Xuankun Li * [xuankun.li@csiro.au]; David K. Yeates [david.yeates@csiro.au] — 2 Research School of Biology, Australian National University, Canberra, ACT 2601, Australia — * Corresponding author
Electronic Supplement: Additional material examined.
Eristalopsis australis (Yeates)
Australia, NT, ♂ (1), Keep River National Park, 4 km SE Jarrnarm Camp Ground, hand netted;
1–3-vi-2001, FD Parker 15°47′49″S 129°06′31″E (ANIC: 29-038740). ♂ (1), Keep River National Park, Ranger Station, pan trap in grassland; 3-v-2001, FD Parker & ME Irwin, 15°58′19″S 129°02′18″E (ANIC: 29-038741). ♂ (29) ♀ (1), Keep River National Park, Bail-Me- Up Cr 23.7 km SSW Jarrnarm Camp Ground, malaise in dry creek bed; 31-v–3-vi-2001, ME Irwin, FD Parker & C Lambkin, 15°57′55″S 129°01′52″E (ANIC: 29-038742–29-038784, 29- 038516–29-038524 [♂];: 29-038525 [♀]). ♂ (3), Keep River National Park, Bail-Me-Up Cr 23.7 km SSW Jarrnarm Camp Ground, malaise in dry creek bed; 3–8-vi-2001, ME Irwin, FD Parker
& C Lambkin, 15°57′55″S 129°01′52″E (ANIC: 29-038526–29-038528). ♂ (33), Keep River National Park, Bail-Me-Up Cr 23.7 km SSW Jarrnarm Camp Ground, malaise in dry creek bed;
9–10-vi-2001, ME Irwin, FD Parker & C Lambkin, 15°57′55″S 129°01′52″E (ANIC: 29- 038529–29-038561). ♂ (22) ♀ (1), Keep River National Park, Bail-Me-Up Cr 23.7 km SSW Jarrnarm Camp Ground, malaise in dry creek bed; 10–13-vi-2001, ME Irwin, FD Parker & C Lambkin, 15°57′55″S 129°01′52″E (ANIC: 29-038473–29-038494 [♂]; 29-038495 [♀]).
Australia, WA, ♂ (1), 114 km NW Meekatharra, 22-ix-[19]95, James O′Grady (ANIC: 29- 038562). ♀ (3), Gascoyne River at Highway 95; 19-ix-1983, EI Schlinger & ME Irwin, river wash with white-barked eucalyptus (ANIC: 29-038563–29-038565). ♀ (1), 1 km N Jct. Highway 95 and Neds Creek Road; 19-ix-1983, EI Schlinger & ME Irwin, Ptilotis in bloom (ANIC: 29- 038566).
Eristalopsis byrrhus (Bowden)
Australia, WA, ♂ (1), Pinjar, Perry Rd., 31°33′S 115°49′E, 8-xi-1981, MJ Smart (MSC). ♀ (1), Pinjar, Perry Rd., 31°35′S 115°48′E, 30-x-1982, MJ Smart (MSC). ♀ (1), Long Point 9 mi, SW Walpole, 13-x-1970, GA Holloway (AMS: K504553). ♂ (3) ♀ (1), Swan River, Inglewood, BA O′Connor, October (WADA: 72198–72201). ♀ (1), Swan River, Perry, Gnangara, 21-xi-1930 (WADA: 72202). ♀ (1), Swan River, Inglewood, BA O′Connor, 21-xi-1930 (WADA: 72203). ♂ (1) Swan River, LJ Newman, November (WADA: 72204). ♂ (6) ♀ (5), Capel, 10-xi-1940, PNF (WADA: 72204–72216). ♂ (1) Swan River, LJ Newman, October (WADA: 72217).
Eristalopsis concava sp.n.
Australia, NT, ♂ (1), Birraduk Ck., 17 km WSW of Nimbuwah Rock; 4-vi-1973, DH Colless (ANIC: 29-038573). ♀ (1), Birraduk Ck., 11 km WSW of Nimbuwah Rock; 4-vi-1973, DH
Colless (ANIC: 29-038574). ♂ (1), Keep River National Park, Bail-Me-Up Cr 23.7 km SSW Jarrnarm Camp Ground, hand netted; 1/3-vi-2001, FD Parker, 15°47′49″S 129°06′31″E (GPS) (ANIC: 29-038575). ♂ (1), Narbarlek Dam, 15 km S by W of Nimbuwah Rock; 2-vi-1973, MS Upton, RS McInnes (ANIC: 29-038576). ♂ (1), Nourlangie Ck., 8 km N of Mt. Cahill; 17-vi- 1973, DH Colless (ANIC: 29-038577). ♀ (1), Cooper Creek, 11 km S by W of Nimbuwah Rock;
4-vi-1973, DH Colless (ANIC: 29-038578). ♀ (2), Keep River National Park, Bail-Me-Up Cr., 23.7 km SSW Jarrnarm camp ground, dry creek bed hand net; 9-vi-2001, ME Irwin, FD Parker, CL Lambkin, 15°57′55″S 129°01′52″E (GPS) (ANIC: 29-038580–29-038581). Australia, WA,
♀ (1), Calm Site 13/4 12 km S of Kalumburu Mission, open forest; 7–11-vi-1988, TA Weir (ANIC: 29-038579).
Eristalopsis parvi sp.n.
Australia, NT, ♂ (2) ♀ (1), Koongarra area, 15–22 km E of Mt. Cahill, 9-iii-1973, DH Colless (ANIC: 29-041453–29-041454 [♂]; 29-041455 [♀]). ♀ (2), Koongarra, 15 km E of Mt. Cahill, 12-vi-1973, DH Colless (ANIC: 29-041456–29-041457). ♂ (1), Oberie Rock, 21-x-1972, EF Riek (ANIC: 29-041458). ♂ (1), Nawurlandja Rock area, Kakadu National Park, 100 m, 21°51′S 132°47′E, 22-i-1993, D and A Daniels (AMS: K504591). ♂ (1), Katherine Gorge, near Katherine, 15-iii-1973, EM Exley (QM).
Eristalopsis robertsi (Paramonov)
Australia, QLD, ♂ (1) ♀ (1), North Stradbroke Is., SEQLD, 5-iv-1987, DK Yeates (QM). ♂ (1)
♀ (4), North Stradbroke Is., SEQLD, 4-iv-1987, DK Yeates (QM). ♀ (1), North Stradbroke Is., 15-iii-1986, DK Yeates (QM). ♀ (10), Youth hostel, Cape Tribulation, 8–10-x-1984, DK Yeates (QM). ♂ (3) ♀ (5), 7 km E Dunwich, North Stradbroke Island, 120 m, 27°30′S 153°27′E, 27-iii- 1993, G Daniels, CJ Burwell (QM). ♂ (2), 13 km N of Dunwich, North Stradbroke Is., 26°55′S 153°09′E, 5-iv-1987, G Daniels (QM). ♂ (1) ♀ (1), 13 km N of Dunwich, North Stradbroke Is., 26°55′S 153°09′E, 16-iii-1986, DK Yeates (QM). ♀ (1), 13 km N of Dunwich, North Stradbroke Is., 27°26′S 153°27′E, 18-iii-1990, G Daniels (QM). ♀ (2), Shiptons Flat, 37 km S Cooktown, open forest, 29-xi-1985, DK Yeates (QM). ♀ (1), C.R.E.B. track 16.5 km N Daintree, Rainforest, 520 m, 27-xi-1985, DK Yeates (QM). ♂ (1), 3.5 km WSW of Point Lookout, North Stradbroke Is., 27°26′S 153°30′E, 4-iv-1992, G Daniels, CJ Burwell (QM). ♀ (1), Central Station, Fraser Island, 24–28-xi-1977, K Walker (QM). ♂ (1), Mulgrave R. Vly, 14.5 km SbyW of Gordonvale, 16-xi-1981, DH Colless (QM). ♂ (2), rainforest nr Rainbow Beach, 15-xi-1981, G and A Daniels (AMS: K504622–K504623). ♂ (1), Station Creek, 17 km NW of Mount Molloy, 16°37′S 145°13′E, 4-i-1984, G and A Daniels (AMS: K504624). ♂ (1) ♀ (1), Garner′s Beach, nr Bingil Bay, 12-x-1984, NW Rodd (AMS: K504625–K504626). ♀ (1), Mount Perserverance, W of Mossman, 16°33′S 145°20′E, 5-i-1985, G and A Daniels (AMS: K504627). ♂ (1), 3.5 km WSW of Point Lookout, North Stradbroke Is., 27°26′S 153°30′E, 10-iii-1991, G Daniels (AMS:
K504629). ♀ (2), Shiptons Flat 31 km S Cooktown, R. Eastwood rainforest, 15°47′S 145°13′E, 21-i-1994, G and A Daniels (AMS: K504630–K504631). ♀ (1), Garners Beach, 17°49′S 146°06′E, 17-x-1995, R. Eastwood (AMS: K504632). ♀ (1), 19 km West Paluma, 19-i-1980, S Johnson (AMS: K504633). ♂ (1) ♀ (1), Kuranda, North Q, 14-x-1973, G Williams (AMS:
K504634–K504635). ♀ (2), rainforest nr Rainbow Beach, 15-xi-1981, G and A Daniels (AMS:
K504636–K504637). ♀ (1), Great Sandy NP, Bymien Picnic Area, Rain Forest, 25°57′13″S 153°6′16″E, DK Yeates and X Li, 7 Dec 2017 (ANIC: 29-040681).
Eristalopsis rubra sp.n.
Australia, NSW, ♂ (2), 6 km NE Bilpin nr. Kurrajong, 14-ix-1979, NW Rodd (AMS: K504610–
K504611).
Eristalopsis wrightae sp.n.
Australia, QLD ♂ (1) ♀ (5), N Bundaberg, 6-iii-1972, H Frauca (ANIC: 29-041466 [♂]; 29- 041467–29-041471 [♀]). ♂ (1) ♀ (4), Bundaberg, 1–2-x-1971, H Frauca (ANIC: 29-041472 [♂];29-041473–29-041476 [♀]). ♂ (2), Nth. Bundaberg, 30-xi-1971, H Frauca (ANIC: 29- 041477–29-041478). ♂ (1) ♀ (1), Bundaberg, i.1971, H Frauca (ANIC: 29-041479–29-041480).
♂ (2), Arcadia Magnetic Is., 19.09°S 146.52°E, 6-x-1998, RW Matthews (ANIC: 29-041481–29- 041482). ♀ (2), Arcadia Magnetic Is., 19.09°S 146.52°E, 13-x-1998, RW Matthews (ANIC: 29- 041483–29-041484). ♀ (1), Isabella Falls, 15.18°S 145.00°E, 19-i-1994, P Zborowski, RW Matthews (ANIC: 29-041485). ♂ (2) ♀ (3), Kings Plains, Quinkan Bush Blitz Helipad 13., 15.492°S 144.828°E, 14-iii-2017, SG Wright, LW Popple (QM). ♀ (1), Bonnyglen, BushBlitz site B9, 15.868°S 144.653°E, 9-iii-2017, Wright, Popple & Burwell (QM). ♂ (3), Captain Billy Ck x-ing, Cape York Pen., 11°37′S 142°49′E, 11-iii-1992, G Daniels, MA Schneider (QM). ♀ (1), Iron Ra., West Claudie R., 3-x-1985, D Yeates (QM). ♀ (1), Iron Ra., West Claudie R., 8-x- 1985, D Yeates (QM). ♀ (1), Iron Ra., West Claudie R., 9-x-1985, D Yeates (QM). ♂ (1), Central Station, Fraser Island, 24–28-xi-1977, K Walker (QM). ♀ (1), Binna Burra, 7-ix-1944, OW Sturgess (QM). ♀ (1), Noosa Nat. Pk., 13-x-1965, G Monteith (QM). ♀ (1), McIvor Riv.
Rd. 4 km N Isabella Falls, 15°15′S 145°00′E, 300 m, 17-i-1994, G and A Daniels, R Eastwood (AMS: K504548). ♂ (1), Eclectus, Iron Range, 12°45′46″S 143°17′10″E, 20 m, 6-i-1996, G and A Daniels (AMS: K504550). ♂ (1), Carnarvon Rge., 18-x-1938, N Geary (AMS: K504551). ♀ (1), ′Bald Hills′ Stn, 9 km N of Isabella Falls, 15°15′S 145°00′E, 1-i-1985, G and A Daniels (AMS: K504552). ♀ (1), Cooloola N.P. 28–29-xi-1985, D Bickel and G Cassis (AMS:
K504554). ♀ (1), 3 km SW Fox Ck x-ing, ′Wolverton′, 13°11′S 142°56′E, 13-iv-1989, G and A Daniels (AMS: K504555). ♀ (1), Archer River x-ing, 13°25′S 142°56′E, 6-iv-1989, G and A Daniels (AMS: K504556). ♂ (1) ♀ (1), Archer River x-ing, 13°25′S 142°56′E, 7-iv-1989, G and A Daniels (AMS: K504557–K504558). ♀ (1), 17 km S of McIvor Riv., 9-i-1982, G and A Daniels (AMS: K504559). ♀ (1), Mount Cook, nr Cooktown, 12-i-1982, G and A Daniels (AMS: K504560). ♀ (1), ′Bald Hills′ Stn, 8 km N of Isabella Ck x-ing, NW of Cooktown, 15°15′S 145°00′E, 13-i-1982, G and A Daniels (AMS: K504561). ♀ (1), ′Bald Hills′ Stn, 7.6 km N of Isabella Ck x-ing, NW of Cooktown, 15°15′S 145°00′E, 10-i-1982, G and A Daniels (AMS:
K504562). ♀ (1), ′Bald Hills′ Stn, 7.6 km N of Isabella Ck x-ing, NW of Cooktown, 15°15′S 145°00′E, 9-i-1982, G and A Daniels (AMS: K504563). ♀ (1), 3 km NW Archer Riv., x-ing, 13°24′S 142°55′E, 11-iv-1989, G and A Daniels (AMS: K504564). ♀ (1), Archer Riv., x-ing, 13°25′S 142°56′E, 7-iv-1989, G and A Daniels (AMS: K504565). ♀ (1), 1 km N of Eclectus Iron Range, 12°45′45″S 143°17′11″E, 20 m, 6-i-1996, G and A Daniels (AMS: K504566). ♂ (1), Rainbow Beach, 20-xi-1979, G Daniels (AMS: K504567). ♂ (1), Mcllwrath Ra. 8 km NE Coen, 13°53′30″S 143°15′21″E, 540 m, 13-i-1994, G and A Daniels, R Eastwood (AMS: K504568). ♂ (1), Cathu State Forest, 20°47′S 148°32′E, 120 m, G and A Daniels (AMS: K504569). ♀ (1), 13 km W Musgrave, 14°48′S 143°23′E, 26-iv-1989, G and A Daniels (AMS: K50457). ♂ (2), West Claudie Riv. x-ing, Iron Range, 12°44′S 143°13′E, 50 m, 8-i-1994, G and A Daniels, R Eastwood (AMS: K504571–K504572). ♂ (1), Battle Camp Rd., Isabella Falls x-ing, 15°18′05″S 145°00′15″E, 240 m, 3-i-1994, G and A Daniels, R Eastwood (AMS: K504573). ♂ (1), Shiptons Flat, 31 km S Cooktown, 15°47′S 145°13′E, 20-i-1994, G and A Daniels, R Eastwood (AMS:
K504574). ♀ (1), Charleville–Bollon Rd., 70 km SE Charleville, 26°52′S 146°35′E, 300 m, 30- ix-1991, G Daniels (AMS: K504575). ♀ (1), Shiptons Flat, 31 km S Cooktown, 15°47′S 145°13′E, 20-i-1994, G and A Daniels, R Eastwood (AMS: K504576). ♀ (2), Battle Camp Rd., Isabella Falls x-ing, 15°17′S 145°53′E, 200 m, 17-i-1994, G and A Daniels, R Eastwood (AMS:
K504577–K504578). ♂ (1), 12 km N of Palmer River x-ing, 16°01′S 143°49′E, 4-i-1985, G and A Daniels (AMS: K504579). ♀ (1), Battle Camp Rd., 7 km E Normanby Riv., x-ing, 15°17′S 145°54′E, 160 m, 17-i-1994, G and A Daniels, R Eastwood (AMS: K504580). ♀ (3), McIvor Riv. Rd. 4 km N Isabella Falls, 15°15′S 145°00′E, 300 m, 17-i-1994, G and A Daniels, R Eastwood (AMS: K504581–K504583). ♂ (1), ′Bald Hills′ Stn, 9 km N of Isabella Falls, 15°14′S 145°01′E, 29-x-1984, G and A Daniels (AMS: K504584). ♂ (1), 13 km Musgrave, 15°14′S 145°01′E, 26-iv-1989, G and A Daniels (AMS: K504585). ♂ (1), Eclectus, Iron Range, 12°45′46″S 143°17′10″E, 20 m, 26-x-1995, G and A Daniels (AMS: K504586). ♀ (1), Eclectus, Iron Range, 12°45′46″S 143°17′10″E, 20 m, 5-i-1996, G and A Daniels (AMS: K504587). ♂ (1), Clarke Ra., South Rd., Cathu St. For., 20°49′21″S 148°32′30″E, 780 m, G and A Daniels (AMS:
K504588). ♂ (1), Eclectus, Iron Range, 12°45′46″S 143°17′10″E, 20 m, 27-x-1995, G and A Daniels (AMS: K504589). ♂ (1), Eclectus, Iron Range, 12°45′46″S 143°17′10″E, 20 m, 5-i- 1996, G and A Daniels (AMS: K504590). ♂ (2) ♀ (1), Mcllwrath Ra. 8 km NE Coen, 13°53′30″S 143°15′21″E, 14-i-1994, 540 m, G and A Daniels, R Eastwood (AMS: K504592–
K504594). ♀ (1), West Claudie Riv. x-ing, Iron Range, 12°44′S 143°13′E, 9-i-1994, 50 m, G and A Daniels, R Eastwood (AMS: K504595). ♀ (1), 13 km W Musgrave, 14°48′S 143°23′E, 26-iv- 1989, G and A Daniels (AMS: K504596). ♀ (1), Byfield State Forest, 1-i-1976, G Daniels (AMS: K504597). ♀ (1), Wildash nr Warwick, 28°18′S 152°03′E, 500 m, 14-i-1993, R Eastwood (AMS: K504599). ♂ (2) ♀ (1), 20 km S Warwick, 28°20′S 151°56′E, 650 m, 10-x- 1992, R Eastwood (AMS: K504602–K504604). ♂ (1), Mcllwrath Ra. 8 km NE Coen, 13°53′30″S 143°15′21″E, 13-i-1994, 540 m, G and A Daniels, R Eastwood (AMS: K504605). ♂ (1), 9 mile NE Mt. Lamond Iron Range, 15-x-1971, DK McAlpine, GA Holloway, DP Sands (AMS: K504606). ♀ (1), 17 km S of McIvor Riv., 9-i-1982, G and A Daniels (AMS: K504612).
♀ (1), Cape York, 2-vi-1985, NW Rodd (AMS: K504613). ♀ (1), Tinaroo Creek Road, nr Mareeba, 21-i-1982, G and A Daniels (AMS: K504614). ♀ (1), Clar Kson Ck., 11 km S Cardwell, 27-x-1980, G Daniels (AMS: K504615). ♀ (1), 22 km S of McIvor Riv., 9-i-1982, G and A Daniels (AMS: K504616). ♀ (1), Baldy Mountain nr. Atherton, 20-i-1982, G and A Daniels (AMS: K504617). ♂ (1), Rainbow Beach, 20-xi-1979, G Daniels (AMS: K504618). ♂ (1), 17 km S of McIvor Riv., 9-i-1982, G and A Daniels (AMS: K504619). ♀ (1), McLeod River x-ing, nr Mount Carbine, 7-i-1982, G and A Daniels (AMS: K504620). ♀ (1), 12 km N of Palmer River x-ing, 16°01′S 144°49′E, 4-i-1985, G and A Daniels (AMS: K504621). ♀ (4), Great Sandy NP, Bymien Picnic Area, Rain Forest, 25°57′13″S 153°6′16″E, DK Yeates and X Li, 7 Dec 2017 (ANIC: 29-040677–29-040680).
