Neaporia maculata (Weise), new combination

Prodilis maculata Weise 1902: 175; Korschefsky 1931: 109; Blackwelder 1945: 444.

Description. Female holotype. Length 2.5 mm, width 1.8 mm; body elongate oval, elytron with side curved, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny. Color black;

head bluish black; pronotum bluish black except lateral margin narrowly yellow; elytron black with large, yellow, lateral macula in basal 2/3, inner margin of macula curved from anterolateral angle of elytron 3/4 distance to sutural margin medially, then curved outward to lateral margin at apical decliv-ity (Fig. 303); antenna yellow; mouthparts yellow except apical maxillary palpomere with apical 2/3 dark brown; epipleuron, legs yellow; ventral surface yellowish brown. Head punctures small, separated by less than a diameter; pronotal punctures as large as on head, separated by a diameter or less; elytral punctures slightly larger than on pronotum, separated by less than 3 times a diameter; prosternal, mesosternal punctures large, separated by less than a diameter; metasternum with large punctures along anterior and lateral borders, punctures on remaining surface small, sparse; punctures on abdomi-nal ventrites 1–3 large, separated by a diameter or less, punctures on remaining ventrites small, sepa-rated by about a diameter. Head not densely pubescent, frons slightly widened from vertex to clypeus, about 1½ times as wide as eye measured at vertex (Fig. 304); eye canthus short; apical maxillary palpomere long, narrowed to apex in apical 1/3. Pronotum widest at middle, reflexed lateral margin narrow, wid-ened from base to apex. Epipleuron flat, wide in basal ½, as wide as pronotal hypomeron. Prosternum longer than wide, longer than mesosternum. Postcoxal line on ventrite 1 short, narrowly rounded, ex-tended slightly beyond midpoint of ventrite. Apex of ventrite 5 arcuate medially. Genitalia not exam-ined.

Male. Unknown.

Variation. Unknown.

Type locality. PERU: Marcapata Type depository. ZMHB.

Geographical distribution. Peru.

Specimens examined. 1. Holotype.

Remarks. Neaporia maculata is distinguished from other species of Neaporia by the unique dorsal color pattern. The female holotype is labeled “Typus (red paper)/ex coll. J. Weise / marcapata (green

G H paper)/Prodilis maculata w”. It is apparent from Weise’s original description that he had a single type specimen here considered to be a holotype.

Succinctonotum Gordon and Hanley, new genus

Description. Cephaloscymnini with body form short, widely oval, narrowed from apex of pronotum to apex of abdomen. Frons narrow, widened from vertex to apex, widest point nearly as wide as eye; apex of frons not extended beyond antennal insertion; male clypeus and anterior portion of frons pale yellow or yellowish white, not densely pubescent (Fig. 308); female head without maculation. Apical maxillary palpomere slender, narrowed from middle to apex as in Neaporia. Pronotum short, about two times as long as scutellum, with anterior margin deeply excavated for reception of head, with oblique surface groove as in Neaporia, strongly projected forward laterally to midpoint of eye (Fig. 308). Prothoracic hypomeron without fossa. Epipleuron narrow, flat. Male pro–, meso– and metasterna mostly flat except weak depression between pro– and mesosterna; prosternum short, wide, without lateral carina, apex without modified setae. Male metasternum without pit medially adjacent to metepisternum. Tarsal claw without basal angulation. Apex of male 5th ventrite arcuate.

Remarks. This genus is thus far known from two specimens. It is recognized by the short body; pronotum only slightly longer than scutellum medially; narrow, elongate head; and flat, unmodified prosternal process. The male is described below and female genitalia of the other specimen are also described, but the female is not designated as a type because it cannot be determined with certainty that these speci-mens are the same species. Succinctonotum is similar to Neaporia, sharing the same pronotal groove and lack of prosternal carinae.

Etymology. The genus name refers to the short pronotum of the type species; gender feminine.

Succinctonotum frosti Gordon and Hanley, new species

Description. Male holotype. Length 1.4 mm, width 1.2 mm. Dorsal surface entirely shiny, lacking microsculpture. Color black except head reddish yellow (Fig. 308), with black vertex, maculae consisting of narrow, yellow, sinuate vitta from clypeal apex adjacent to eye onto vertex, and short, yellow, median vitta from clypeal apex to midpoint of eye (Fig. 310); pronotum with anterolateral angle reddish yellow;

elytra with brownish red margin; antenna, legs yellow; mouthparts yellow except apical 1/3 of final palpomere brown; prosternal process brownish red; remainder of ventral surface brown with reddish tint except ventrite 5 yellow. Head punctures coarse, separated by a diameter or less; pronotal punc-tures smaller than head puncpunc-tures, separated by a diameter or less; elytral puncpunc-tures slightly larger than pronotal punctures, separated by a diameter or twice a diameter; prosternal, metasternal punc-tures large, separated by less than a diameter, puncpunc-tures on abdominal ventrites 1, 2 large, separated by a diameter or less, punctures on remaining ventrites small, separated by about a diameter. Head with frons slightly more than ½ width of eye measured at vertex; eye canthus short (Fig. 310); apical maxil-lary palpomere long, narrowed to apex in apical ½. Pronotum widest at middle, reflexed lateral margin narrow. Epipleuron flat, narrow. Prosternum short, as wide as long, as long as mesosternum, with apical margin nearly truncate, small anterolateral projection not setose. Postcoxal line on ventrite 1 short, arcuate, extended ½ distance to apical margin of ventrite (Fig. 309). Apex of ventrite 5 narrowly, shal-lowly emarginate medially. Genitalia with basal lobe longer than paramere, slender, about equal in width from base to deeply emarginate apex; paramere slender, slightly narrowed from base to acute apex, weakly sinuate, dorsal margin smooth (Fig. 311, 312); sipho lost.

Female. Similar to male except head not maculate, black, elytron bluish black (Fig. 314); Spermathecal capsule curved medially, base enlarged, remainder of capsule slender (Fig. 313).

Variation. Unknown.

Type material. Holotype male; PANAMA: BarrowColoIsl. (Barrow Colorado Island, CZ (Canal Zone), Feb. 11–1929. (USNM).

Other specimen. 1. Female differing from male described above by length 1.3 mm, width 1.1 mm;

immaculate head; distinctly blue elytra; and prosternal process medially depressed. Genitalia typical of Cephaloscymnini, spermathecal capsule lost. Label data: COSTA RICA: Heredia Est. Biol. La Selva, 50–150m 10o 26'N 84o01'W, Prov. ALAS, INBIO–OET, M/06/021, 02 Marzo 1983, Bosque primario.

(USNM).

Remarks. These specimens are distinguished from other known Cephaloscymnini by the generic char-acters. The female listed under “other specimen” above is not designated as a type because it may not be conspecific with the male holotype. It matches the holotype in all characters other than those described above.

Etymology. This species is named for S. W. Frost, collector of the holotype.

Prodilis Mulsant

Prodilis Mulsant 1850: 898; Crotch 1874: 276; Weise 1904: 363 (Neaporia placed as a synonym of Prodilis);

Korschefsky 1931: 109; Blackwelder 1945: 444. Type species. Prodilis pallidifrons Mulsant, 1850, by monotypy.

Scymnus (Volgus) Mulsant 1853: 147: Korschefsky 1931: 116. NEW SYNONYM. Type species. Scymnus (Polius) volgus Mulsant 1853, by monotypy.

Prodiloides Weise 1922: 36; Korschefsky 1931: 110. NEW SYNONYM. Type species. Prodiloides bipunctata Weise 1922, by monotypy

Aneaporia Casey 1908: 407; Korschefsky 1931: 108. Type species. Neaporia plagioderina Gorham 1897, by monotypy.

Description. Cephaloscymnini with body form widely oval to narrowly oval, widest at middle of elytra.

Frons wide, inner margins of eyes parallel or nearly so, usually about twice width of an eye; apex of frons extended beyond antennal insertion usually by about width of basal antennal article except 4 species with elongate frons extended by about 3 times width of basal antennal article; male clypeus and anterior portion of frons usually pale yellow or yellowish white, not densely pubescent (Fig. 329); female head without maculation. Apical maxillary palpomere widened from base to apex (Fig. 306, 307). Pronotum long, with anterior margin moderately excavated for reception of head, weakly projected forward later-ally to about apical 4/5 of eye, pronotum without surface groove (Fig. 315). Prothoracic hypomeron with or without fossa. Epipleuron usually wide, sometimes flat, sometimes with outer margin weakly or strongly descending, occasionally with feeble depressions for reception of femoral apices (P. sabrina type). Male pro–, meso– and metasterna flat or rounded, rarely medially depressed; prosternum not widely expanded to conceal mouthparts or sometimes expanded to completely conceal mouthparts, prosternal process with base truncate or rounded, process wide, long, lateral carina present on each side adjacent to procoxa; male prosternal process without modified setae. Male metasternum without pit medially adjacent to metepisternum. Tarsal claw without basal angulation. Apex of male 5th ventrite arcuate.

Remarks. Polius Mulsant (1853) is monotypic and was described as a subgenus of Scymnus Kugelann.

It is a member of Cephaloscymnini here placed as a synonym of Prodilis. It differs from typical Prodilis by an epipleuron often slightly descending externally. However, this is not consistent because many Polius volgus specimens have an absolutely flat epipleuron. Casey (1908) considered those specimens with a descending epipleuron to belong to Aneaporia, which was considered a subgenus of Prodilis by Korschefsky (1931). Prodiloides Weise is here considered a synonym of Prodilis because the principal defining character, a prosternum long in front, concealing mouthparts in typical Prodiloides and short,

G H