Combined analysis - The Phylogenetic Position of the Isopoda in the Peracarida (Crustacea: Mala

2.4.1. Weighting strategy

The costs (= tree lengths) of the two datasets (18S;

morphology) are substantially different. Morpho logy (202 static characters) contributed approximately 1.5% to the cost of a combined unweighted dataset.

Several higher weightings of the morphology rela-tive to the molecular data were tried, but the equally weighted data are reported here. In the current case, the sequence data matrix used a transformation cost matrix that gives substitutions a cost of 2, indels a cost of 1 and gap opening events a cost of 1, as determined by the monophyly survey (Tab. 2). Equal weightings for all transformations were achieved by assigning a weight of 0.5 to the molecular matrix, and a weight of 1 to all morphological characters. Down weighting the molecular matrix was preferred because the weight-ing is applied to all dynamic transitions equally. Two other possible approaches to equal weights were not employed in this analysis. Under the paradigm that each dynamic homology character is a fragment that transforms during evolution, the dynamic characters could be down weighted so that their contribution would equal to that of each of the static morphologi-cal characters. A second approach to equal weighting would down weight the dynamic characters so that their contribution to the cost of the tree exactly equals that of the static characters. Although these alterna-tive weightings were not tried for this analysis, both might result in the dynamic molecular characters be-ing swamped by the static morphological characters, so that such analyses might resemble the morphologi-cal tree only.

2.4.2. Analyses

The 18S data were analysed as 10 fragments as ex-plained in section 2.2. The morphological data were converted to a Hennig86/Nona format (see G^OLOBOFF et al. 2008 and references cited therein) for input into POY 4. The best dynamic parameter set (substitu-tions = 2, indels = 1, gap opening = 1; see Tab. 2) and weights as explained above was used to analyse 18S and morphological data simultaneously. The relevant script for the iterated combined analysis runs is shown in Tab. 5. Bremer support was calculated in POY 4.1.1 using a script described in V^ARÓN et al. (2008).

Uropod exopod: state 0, present; state 1, absent.

196.

In isopods, the smaller exopod is considered absent if only one ramus is present.

Uropod exopod segments: state 0, one article;

197.

state 1, two articles. No peracarid is known to have more than 2 articles in the uropodal exopod.

Uropod exopod ramus (shape): state 0, rod-like, 198.

rounded tip; state 1, robust, spine-like; state 2, fl at-tened.

Uropod endopod (present/absent): state 0, pre-199.

sent; state 1, absent.

Uropod endopod proximal articulation: state 0, 200.

present; state 1, absent.

Uropod endopod segments (

201. B^RUSCA & W^ILSON

1991: chr 59): state 0, two articles; state 1, one article;

state 2, three or more articles. State 2 is seen in Cuma-cea, Tanaidacea and Hirsutiidae.

202. Uropod endopod ramus (shape) (BRUSCA & WIL

-SON 1991: chr 57): state 0, rod-like, rounded tip; state 1, robust, spine-like; state 2, fl attened.

2.3.4. Morphological data analysis

The morphological data were analysed using PAUP* version 4.0b10 (SWOFFORD 1998), with all character transitions set to unordered, FITCH (1971) parsimony. Starting trees were obtained via random stepwise addition. A treespace sampling protocol (J^UST & W^ILSON 2004) was used, including 10,000 samples of 3 trees (PAUP* commands: hsearch add-seq=random nchuck=3 chuckscore=1 nreps=10000 randomize=trees; hsearch start=current nchuck=0 chuckscore=0). A strict consensus was obtained using PAUP* to assess congruence of the phylogenetic hy-potheses. To assess variation in their topologies, the most distant tree from an arbitrary tree (tree 1) was found using PAUP* symmetric distance tree metric in the treedist and fi lter trees commands. One of the most distant trees found was then used as the root to fi nd other most distant trees. These trees were cho-sen for comparison. Jackknife (F^ARRIS et al. 1996) branch support values were obtained for the com-bined dataset based on 1000 randomised samples of the data (PAUP* settings for each iteration: hsearch addseq=random nchuck=10 chuckscore=1 nreps=10 randomize=trees). The jackknife method used 33%

characters deleted from each of the 1000 replicates.

Bremer support was calculated using a script gener-ated by MacClade 4.0 (M^ADDISON & M^ADDISION 2000) and analysed in PAUP*.

Tab. 4. Morphological data matrix of 75 taxa listed in Tab. 1 and 202 morphological characters. Polymorphism is indicated as fol-lows: A = states 0 and 1; B = states 0 and 2; C = states 1 and 2. See section 2.3.3. for character analysis.

Character # 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 3 3 3 3 3 3 3 3 3 3 4 4 4 4 4 4 4 4 4 4 5 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 Gonodactylus viridis 0 0 0 0 0 0 - 0 - - - 1 - - - - 0 1 1 0 1 - 1 1 0 0 1 0 0 0 0 0 0 1 - 1 0 ? 0 0 0 1 0 0 0 ? ? 0 Penaeus semisulcatus 0 0 1 0 0 0 - 0 - - - 0 - - - - 0 0 1 0 1 - 0 1 0 1 0 0 - 0 0 0 0 0 - 1 0 0 0 0 2 1 0 0 0 ? ? 0 Euphausia superba 0 0 1 0 3 - - 0 - - - 0 - - - - 0 0 0 0 1 - 0 1 0 1 0 0 - 0 0 0 0 0 - 1 0 0 0 0 0 0 - 0 0 ? ? 0 Anaspides tasmaniae 2 ? ? 0 3 - - 0 - - - 0 - - - - 0 0 0 0 0 0 1 1 0 0 1 0 0 0 2 - 0 0 - 1 0 0 0 2 0 0 - 0 0 ? ? 0 Neognathophausia ingens 1 1 0 0 0 0 - 1 0 1 0 1 0 0 - - - - 0 1 0 1 0 1 1 1 0 0 0 0 - 0 0 1 0 0 - 1 0 0 0 0 0 1 0 0 0 ? ? 0 Heteromysis formosa 1 1 0 0 0 0 - 1 0 1 0 1 0 0 - - - - 0 1 0 1 0 1 1 1 0 0 0 0 - 0 1 1 0 0 - 1 0 0 0 0 2 1 1 0 0 ? ? 0 Stygiomysis holthuisi 1 1 0 0 0 0 - 1 0 1 0 1 0 0 - - - - 0 1 0 1 0 1 1 1 0 0 0 0 - 0 0 1 0 0 - 1 0 0 0 0 2 1 1 0 0 ? ? 0 Thetispelecaris remex ? ? ? 1 3 - 0 1 B 1 0 0 0 ? - - - - 1 ? ? ? ? ? 1 ? 1 0 ? ? ? ? ? ? ? ? ? ? ? ? ? 2 0 ? ? ? ? ? ? 1 Ingolfi ella tabularis 1 1 0 0 3 0 2 1 0 1 0 0 0 0 - - - - 0 0 1 - 0 0 1 1 0 0 0 0 - 0 0 0 0 3 - - 0 0 0 0 2 1 1 0 0 ? ? 0 Arrhis phyllonyx 1 1 0 0 3 0 2 1 0 1 0 0 0 0 - - - - 0 0 1 - 0 0 1 1 0 0 0 0 - 0 0 0 0 3 - - 0 0 0 0 2 1 1 0 0 ? ? 0 Protella gracilis 1 1 0 0 3 0 2 1 0 1 0 0 0 0 - - - - 0 0 1 - 0 0 1 1 0 0 0 0 - 0 0 0 0 3 - - 0 0 0 0 2 1 1 0 0 ? ? 0 Hyperietta stephenseni 1 1 0 0 3 0 2 1 0 1 0 0 0 0 - - - - 0 0 1 - 0 0 1 1 0 0 0 0 - 0 0 0 0 3 - - 0 0 0 0 2 1 1 0 0 ? ? 0 Gammarus troglophilus 1 1 0 0 3 - 2 1 0 1 0 0 0 0 - - - - 0 0 1 - 0 0 1 1 0 0 0 0 - 0 0 0 0 3 - - 0 0 0 0 2 1 1 0 0 ? ? 0 Tethysbaena argentarii 1 1 0 0 0 0 - 0 1 - - - - ? - - - - 0 2 0 0 0 0 1 1 1 ? 0 0 - 0 2 - 1 - - - 1 1 0 0 1 1 1 1 0 ? ? 0 Mictocaris halope 1 1 0 0 0 - 0 1 0 1 0 0 0 0 - - - - 0 ? ? ? ? ? 1 0 1 0 0 0 - 0 2 - 1 - - - 0 1 ? ? ? 1 2 1 1 ? ? 0 Spilocuma salomani 1 1 0 0 0 0 - 1 0 1 0 0 0 0 - - - - 0 1 0 1 0 1 1 0 1 0 0 0 - 0 2 - 0 1 1 - 0 1 0 2 0 1 2 1 0 ? ? 0 Diastylis sculpta 1 1 0 0 0 0 - 1 0 1 1 0 0 0 - - - - 0 1 0 1 0 1 1 0 1 0 0 0 - 0 2 - 0 1 1 - 0 1 0 2 0 1 2 1 0 ? ? 0 Spelaeogriphus lepidops 1 1 0 0 0 1 - 1 0 0 0 0 0 0 - - - - 0 1 ? 1 ? ? ? ? 0 0 0 0 - 0 1 0 1 - - - 1 1 0 2 1 1 2 1 0 ? ? 0 Apseudes bermudeus 1 1 0 1 3 - 2 1 2 1 0 0 0 0 - - - - 1 2 0 1 1 - 1 0 1 0 0 0 - 0 1 - 0 1 1 - 1 1 0 2 0 1 2 1 0 ? ? 0 Kalliapseudes sp. 1 1 0 1 3 - 2 1 2 1 0 0 0 0 - - - - 1 2 0 1 1 - 1 0 1 0 0 0 - 0 1 - 0 1 1 - 1 1 0 2 0 1 2 1 0 ? ? 0 Leptochelia sp. 1 1 0 1 3 - 2 1 2 1 0 0 0 0 - - - - 1 2 0 1 1 - 1 0 1 0 0 0 - 0 1 - 0 1 1 - 1 1 0 1 0 1 2 1 0 ? ? 0 Paratanais malignus 1 1 0 1 3 - 2 1 2 1 0 0 0 0 - - - - 1 2 0 1 1 - 1 0 1 0 0 0 - 0 1 - 0 1 1 - 1 1 0 1 0 1 2 1 0 ? ? 0 Tanais dulongii 1 1 0 1 3 - 2 1 2 1 0 0 0 0 - - - - 1 2 0 1 1 - 1 0 1 0 0 0 - 0 1 - 0 1 1 - 1 1 0 1 0 1 2 1 0 ? ? 0 Paramphisopus palustris 1 1 0 0 0 1 - 1 0 0 1 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 0 0 1 1 0 0 1 0 - 0 1 1 1 0 1 2 1 0 0 0 0 Colubotelson thomsoni 1 1 0 0 0 1 - 1 0 0 1 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 0 0 1 1 0 0 1 0 - 0 1 1 1 0 1 2 1 0 0 0 0 Stenasellus racovitzai 1 1 0 0 0 1 - 1 0 0 1 0 0 1 0 2 1 0 1 1 0 1 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 0 0 1 1 1 0 1 2 1 0 0 0 1 Caecidotea racovitzai 1 1 0 0 3 - 0 1 0 0 1 0 0 1 0 1 1 0 1 1 0 1 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 0 0 1 1 1 0 1 2 1 0 0 0 0 Asellus aquaticus 1 1 0 0 3 - 0 1 0 0 1 0 0 1 0 1 1 0 1 1 0 1 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 0 0 1 1 1 0 1 2 1 0 0 0 0 Stenetriid sp. 1 1 0 0 3 - 0 1 0 1 1 0 0 1 0 1 1 0 3 1 0 1 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 ? 0 1 1 1 0 1 2 1 0 0 0 0 Janira maculosa 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 1 1 0 1 0 1 1 0 1 0 1 2 1 1 1 0 0 1 0 0 0 1 1 1 0 1 2 1 0 ? ? 0 Joeropsis coralicola 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 0 1 0 1 2 1 1 1 0 0 1 0 ? 0 1 1 1 0 1 2 1 0 ? ? 0 Iathrippa trilobatus 1 1 0 0 3 - 2 1 0 0 1 0 0 1 1 4 1 1 1 1 0 1 0 1 1 0 1 0 1 2 1 1 1 0 0 1 0 0 0 1 1 1 0 1 2 1 0 ? ? 0 Haploniscus nudifrons 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Acanthaspidia drygalskii 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Neojaera antarctica 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 0 Thylakogaster sp. 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Dendromunna sp. 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Eurycope sarsi 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Ilyarachna antarctica 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Munnopsis typica 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Betamorpha fusiformis 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Mesosignum cf. usheri 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Macrostylis sp.1 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Janirella sp. 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Ischnomesus sp. 1 1 0 0 3 - 2 1 0 1 1 0 0 1 2 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Eugerdella natator 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Eugerda sp. 1 1 0 0 3 - 2 1 0 1 1 0 0 1 1 4 0 1 2 1 0 1 0 1 1 1 1 0 1 2 1 1 1 0 0 ? 0 ? 0 1 1 1 0 1 2 1 0 ? ? 1 Zonophryxus quinquedens 1 1 0 0 3 - 2 1 0 0 0 0 0 1 - 0 - - 0 0 0 1 0 1 1 0 1 0 1 2 1 1 1 0 1 - - - - 1 1 1 0 1 ? ? 5 ? ? 1 Probopyrus pacifi ciensis 1 1 0 0 3 - 2 1 0 0 0 1 0 1 - 0 - - 0 0 0 1 0 1 1 0 1 0 1 2 1 1 1 0 1 - - - 0 1 1 1 0 1 2 1 5 ? ? 0 Hemiarthrus abdominalis 1 1 0 0 3 - 2 1 0 0 0 1 0 1 - 0 - - 0 0 0 1 0 1 1 0 1 0 1 2 1 1 1 0 1 - - - 0 1 1 1 0 1 2 1 5 ? ? 0 Paragnathia formica 1 1 0 0 3 - 2 1 0 1 - 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 2 0 1 C - 0 ? ? ? 0 1 1 1 0 1 2 1 ? ? ? 0 Riggia paranensis 1 1 0 0 3 - 2 1 0 1 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 2 0 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Anilocra physodes 1 1 0 0 3 - 2 1 0 1 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 2 - 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Cymodoce tattersalli 1 1 0 0 3 - 2 1 0 1 1 0 1 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 1 0 0 2 - 0 ? 1 1 1 0 1 2 1 3 1 1 0 Campecopea hirsuta 1 1 1 0 3 - 2 1 0 1 1 0 1 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 1 0 0 2 - 0 ? 1 1 1 0 1 2 1 1 1 1 0 Sphaeroma serratum 1 1 0 0 3 - 2 1 0 1 1 0 1 1 - 0 - - 0 1 0 1 0 1 0 0 0 0 1 1 0 1 1 0 0 2 0 0 ? 1 1 1 0 1 2 0 3 1 1 0 Cassidinidea sp. 1 1 0 0 3 - 2 0 0 1 1 0 1 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 1 0 0 2 - 0 ? 1 1 1 0 1 2 1 1 1 1 0 Limnoria quadripunctata 1 1 0 0 3 - 0 1 0 ? 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 C - 0 1 0 ? 0 1 1 1 0 1 2 1 1 1 1 0 Eurydice pulchra 1 1 0 0 3 - 2 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 1 0 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Excorallana quadricornis 1 1 0 0 3 - 2 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 1 0 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Natatolana albinota 1 1 0 0 3 - 2 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 1 0 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Aega antarctica 1 1 0 0 3 - 2 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 2 - 0 2 - 0 0 1 1 1 0 1 2 1 3 1 1 0 Ligia oceanica 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Ligia italica 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Ligidium germanicum 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Haplophthalmus danicus 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 1 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Porcellio scaber 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 1 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Oniscus asellus 1 1 0 0 3 - 3 1 0 1 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 1 1 0 1 1 0 1 2 - 0 1 0 - 1 1 1 1 0 1 2 1 1 0 0 0 Paranthura nigropunctata 1 1 0 0 3 - 2 1 0 1 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 2 0 1 1 0 0 3 - - 0 1 1 1 0 1 2 1 4 1 1 0 Cyathura carinata 1 1 0 0 3 - 2 1 0 1 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 0 0 1 2 0 1 1 0 0 3 - - 0 1 1 1 0 1 2 1 4 1 1 0 Pygolabis humphreysi 1 1 0 0 2 - 0 1 0 0 0 0 0 1 - 0 - - 0 1 0 1 0 1 1 0 1 0 1 2 0 1 C - 0 ? ? ? 0 1 1 1 0 1 2 1 1 ? ? 1 Idotea baltica 1 1 0 0 4 - 3 1 0 0 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 1 0 1 1 0 0 1 0 0 1 1 1 1 0 1 2 1 1 1 1 0 Glyptonotus antarcticus 1 1 0 0 4 - 3 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 1 1 0 1 1 0 1 1 0 0 1 0 0 1 1 1 1 0 1 2 1 1 1 1 0 Cleantis prismatica 1 1 0 0 4 - 3 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 1 1 0 1 1 0 1 1 0 0 1 0 0 1 1 1 1 0 1 2 1 1 1 1 0 Antarcturus spinacoronatus 1 1 0 0 4 - 3 1 0 ? 0 0 0 1 - 0 - - 1 1 0 1 0 1 1 0 1 0 1 1 0 1 1 0 0 1 0 ? 1 1 1 1 0 1 2 ? 1 1 1 0

Tab. 4. Continuation.

Character # 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 5 5 5 5 5 5 5 5 5 6 6 6 6 6 6 6 6 6 6 7 7 7 7 7 7 7 7 7 7 8 8 8 8 8 8 8 8 8 8 9 9 9 9 9 9 9 9 9 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 Gonodactylus viridis 0 0 1 0 1 1 0 3 1 4 0 0 0 0 0 0 3 1 0 1 0 0 0 - 0 ? 0 0 0 0 1 0 0 0 0 0 - - 0 0 0 A 0 0 0 0 0 3 0 0 Penaeus semisulcatus 0 0 0 0 0 1 0 3 1 2 0 0 0 0 0 0 3 1 0 0 0 0 0 - 0 1 0 0 0 0 1 1 1 0 0 0 - - 0 0 0 0 0 0 0 0 0 3 1 1 Euphausia superba 0 0 0 1 1 1 0 3 1 2 0 0 0 0 0 0 3 0 0 0 0 0 0 - 0 1 0 0 0 0 1 1 1 0 0 0 - 0 0 0 0 0 0 0 1 0 0 3 0 3 Anaspides tasmaniae 0 0 0 1 0 0 0 0 0 - 2 - 0 0 0 0 3 1 0 0 0 A 1 - 0 1 0 0 1 0 0 1 0 0 0 0 - 0 0 0 0 0 0 0 1 0 0 3 0 3 Neognathophausia ingens 0 0 0 1 1 1 1 2 1 2 1 0 0 0 0 0 3 0 0 0 0 1 1 1 0 1 0 0 1 0 1 1 1 0 0 0 - 0 0 0 0 0 0 1 1 1 0 3 1 2 Heteromysis formosa 0 0 0 1 1 1 1 2 0 - 1 0 0 0 0 0 3 0 0 0 0 1 1 1 0 1 0 0 1 0 1 1 1 0 0 0 - 0 0 0 0 0 0 1 1 1 0 3 0 2 Stygiomysis holthuisi 1 0 - - - - 1 2 1 2 1 0 0 0 0 0 0 2 1 5 0 1 1 0 0 1 0 0 1 0 1 0 1 0 0 0 - 1 0 0 0 0 0 1 1 1 0 3 1 3 Thetispelecaris remex - - - 2 1 1 1 1 0 0 0 0 0 0 1 0 4 1 1 0 0 0 1 0 1 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Ingolfi ella tabularis 1 0 - - - 1 1 1 2 - 0 0 0 0 2 1 1 4 0 1 1 2 1 1 0 0 0 0 0 1 0 0 0 0 - 2 0 2 0 0 2 0 1 1 0 3 1 1 Arrhis phyllonyx 2 0 1 1 1 2 1 1 1 1 2 - 0 0 0 0 2 1 1 0 0 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 - 2 0 2 0 0 2 0 1 1 2 3 1 2 Protella gracilis 2 0 1 1 1 2 1 1 0 - 2 - 0 0 0 0 2 1 1 0 0 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 - 2 0 2 0 0 2 0 1 1 2 3 1 2 Hyperietta stephenseni 2 0 1 1 1 2 1 1 1 1 2 - 0 0 0 0 2 1 1 0 0 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 - 2 0 2 0 0 2 0 1 1 2 3 1 2 Gammarus troglophilus 2 0 1 1 1 2 1 1 1 1 2 - 0 0 0 0 2 1 1 0 0 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 - 2 0 2 0 0 2 0 1 1 2 3 1 2 Tethysbaena argentarii 1 0 - - - 2 0 - 1 2 0 0 0 0 2 1 1 4 4 1 1 1 0 0 0 0 0 0 1 0 1 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Mictocaris halope 1 0 - - - 1 0 - 1 3 0 0 0 0 0 0 1 0 4 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Spilocuma salomani 3 - 1 1 1 3 1 3 0 - 1 2 0 0 0 0 2 1 1 0 4 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 - - 1 1 0 0 0 0 1 1 0 3 0 0 Diastylis sculpta 3 - 1 1 1 3 1 3 0 - 1 2 0 0 0 0 2 1 1 0 4 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 - - 1 1 0 0 0 0 1 1 0 3 0 0 Spelaeogriphus lepidops 1 0 - - - 2 1 1 1 1 0 0 0 0 0 1 1 0 4 1 1 1 0 0 0 0 1 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Apseudes bermudeus 1 0 1 1 1 0 1 2 1 1 1 1 0 0 0 0 0 0 1 0 0 1 1 1 0 0 1 0 0 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Kalliapseudes sp. 1 0 1 1 1 0 1 2 1 1 1 1 0 0 0 0 0 0 1 0 0 1 1 1 0 0 1 0 0 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 0 1 Leptochelia sp. 1 0 1 1 1 0 1 2 1 1 1 1 2 1 1 0 2 0 1 0 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 1 1 Paratanais malignus 1 0 1 1 1 0 1 2 1 1 1 1 2 1 1 0 2 0 1 0 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 1 1 Tanais dulongii 1 0 1 1 1 0 1 2 1 1 1 1 2 1 1 0 2 0 1 0 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 - 0 1 1 0 0 0 0 1 1 0 3 1 1 Paramphisopus palustris 2 0 1 1 1 0 1 1 0 - 2 - 0 0 1 0 2 0 1 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 0 - 2 1 1 1 1 0 0 1 1 0 3 0 1 Colubotelson thomsoni 2 0 1 1 1 0 1 1 0 - 2 - 0 0 1 0 2 0 1 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 0 - 2 1 1 1 1 0 0 1 1 0 3 0 1 Stenasellus racovitzai - - - 1 1 0 - 2 - 0 0 1 0 0 0 0 3 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Caecidotea racovitzai 2 1 1 1 1 0 1 1 0 - 2 - 0 0 1 0 2 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Asellus aquaticus 2 1 1 1 1 0 1 1 0 - 2 - 0 0 1 0 2 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Stenetriid sp. 2 1 1 1 1 0 1 1 1 4 2 - 0 0 1 0 0 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Janira maculosa 2 1 1 1 1 0 1 1 0 - 2 - 0 0 1 0 0 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Joeropsis coralicola 2 1 1 1 1 0 1 1 1 4 2 - 2 1 1 0 2 0 0 4 0 1 1 0 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Iathrippa trilobatus 2 1 1 1 1 0 1 1 1 2 2 - 0 0 1 0 0 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Haploniscus nudifrons - - - 1 1 4 2 - 2 1 1 0 1 0 0 0 0 1 1 2 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Acanthaspidia drygalskii - - - 1 1 2 2 - 0 0 1 0 1 0 0 4 0 1 1 0 0 1 0 0 1 0 0 0 0 1 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Neojaera antarctica 2 1 1 1 1 0 1 2 0 - 2 - 2 1 1 0 0 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Thylakogaster sp. - - - 1 0 - 2 - 0 0 1 0 2 0 0 4 0 1 1 0 0 1 0 0 1 0 0 0 0 0 0 0 - 2 1 1 1 1 0 0 1 1 0 3 0 1 Dendromunna sp. - - - 1 0 - 2 - 0 0 1 0 2 0 0 0 0 1 1 0 0 1 0 0 1 0 0 0 0 0 0 0 - 2 1 1 1 1 0 0 1 1 0 3 0 1 Eurycope sarsi - - - 1 0 - 2 - 0 0 1 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Ilyarachna antarctica - - - 1 0 - 2 - 0 0 1 0 0 0 0 5 1 1 1 0 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Munnopsis typica - - - 1 0 - 2 - 0 0 1 0 0 0 0 5 1 1 1 0 0 1 0 0 1 0 0 0 0 0 0 0 - 2 1 1 1 1 0 0 1 1 0 3 0 1 Betamorpha fusiformis - - - 1 0 - 2 - 0 0 1 0 0 0 0 0 0 1 1 0 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Mesosignum cf. usheri - - - 1 1 4 2 - 0 0 1 0 2 0 0 4 0 1 1 2 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Macrostylis sp.1 - - - 1 1 4 2 - 2 1 1 0 2 0 0 3 0 1 1 2 1 1 0 0 1 0 0 0 0 0 1 1 0 2 1 1 1 1 0 0 1 1 0 3 0 1 Janirella sp. - - - 1 1 2 2 - 0 0 1 0 0 0 0 0 0 1 1 0 0 1 0 0 1 0 0 0 0 1 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Ischnomesus sp. - - - 1 1 4 2 - 0 0 1 0 2 0 0 0 0 1 1 0 A 1 0 0 1 0 0 0 0 1 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Eugerdella natator - - - 1 1 4 2 - 2 1 1 0 0 0 0 3 0 1 1 0 0 1 0 0 1 0 0 0 0 0 1 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Eugerda sp. - - - 1 1 4 2 - 2 1 1 0 0 0 0 3 0 1 1 0 0 1 0 0 1 0 0 0 0 0 0 1 1 2 1 1 1 1 0 0 1 1 0 3 0 1 Zonophryxus quinquedens - - - - 1 - - 1 0 - 2 - 0 1 1 1 2 2 1 4 3 0 0 - 1 0 0 1 1 0 0 4 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 1 1 Probopyrus pacifi ciensis 2 2 1 1 1 0 1 1 0 - 2 - 0 1 1 1 2 2 1 4 3 0 0 - 1 0 0 1 1 0 0 4 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Hemiarthrus abdominalis 2 2 1 1 1 0 1 1 0 - 2 - 0 1 1 1 2 2 1 4 3 0 0 - 1 0 0 1 1 0 0 4 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Paragnathia formica 2 1 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 4 3 0 0 - 1 0 0 1 1 0 0 2 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Riggia paranensis 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 3 0 0 - 0 0 0 0 1 1 0 2 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 1 1 Anilocra physodes 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 3 0 0 - 0 0 0 0 1 1 0 2 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 1 1 Cymodoce tattersalli 2 3 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 0 0 1 1 1 0 1 0 0 1 1 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 0 1 Campecopea hirsuta 2 3 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 0 0 1 1 1 1 ? 0 0 0 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 0 1 Sphaeroma serratum 2 3 0 1 1 0 1 1 0 - 0 - 0 0 1 1 2 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 0 1 Cassidinidea sp. 2 3 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 0 0 1 1 1 0 1 0 0 1 1 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 0 0 1 Limnoria quadripunctata 2 1 1 1 1 0 1 1 0 - 2 - 2 1 1 1 2 0 1 5 2 1 1 1 0 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 1 0 1 Eurydice pulchra 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 0 1 1 1 0 1 0 0 1 0 0 1 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 0 1 1 Excorallana quadricornis 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 3 0 0 - 0 1 0 0 1 1 0 1 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 0 1 1 Natatolana albinota 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 0 1 1 1 0 1 0 0 1 0 0 1 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 0 1 1 Aega antarctica 2 2 1 1 1 0 1 1 0 - 2 - 0 0 1 1 2 0 1 1 3 0 0 - 0 1 0 0 1 1 0 1 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 0 1 0 Ligia oceanica 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Ligia italica 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Ligidium germanicum 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Haplophthalmus danicus 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Porcellio scaber 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 2 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Oniscus asellus 2 1 1 1 1 0 1 1 0 - 2 - 1 1 1 0 2 0 1 2 0 1 1 1 1 1 0 0 1 0 0 3 0 0 0 0 - 0 1 1 1 1 1 0 1 - 1 2 0 1 Paranthura nigropunctata 2 1 1 1 1 0 1 1 0 - 2 - 0 1 1 0 2 0 1 1 3 2 1 1 0 1 0 0 1 0 0 4 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 1 1 1 Cyathura carinata 2 1 1 1 1 0 1 1 0 - 2 - 0 1 1 0 2 0 1 1 0 2 1 1 0 1 0 0 1 0 0 4 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 1 1 1 Pygolabis humphreysi - - - - 1 - - 1 0 - 2 - 0 0 1 0 2 0 0 0 0 1 1 1 0 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 2 0 1 1 1 1 0 1 Idotea baltica 2 1 1 1 1 0 1 1 0 - 2 - 2 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Glyptonotus antarcticus 2 1 1 1 1 0 1 1 0 - 2 - 2 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Cleantis prismatica 2 1 1 1 1 0 1 1 0 - 2 - 2 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1 Antarcturus spinacoronatus 2 1 1 1 1 0 1 1 0 - 2 - 2 1 1 0 2 0 1 0 0 1 1 1 1 1 0 0 1 0 0 0 0 0 0 0 - 1 1 1 1 1 1 0 1 - 1 2 0 1

Tab. 4. Continuation.

Character # 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 3 3 3 3 3 3 3 3 3 3 4 4 4 4 4 4 4 4 4 4 5 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 Gonodactylus viridis 0 2 - 1 0 1 0 - 2 1 0 - - - 1 0 2 0 0 0 0 0 - - - - 0 0 0 1 - 0 0 0 0 0 1 0 - 0 0 0 0 0 0 0 0 3 Penaeus semisulcatus 0 0 0 1 0 1 0 - 2 1 0 - - - 1 - - 0 - 0 0 0 0 0 2 2 2 1 - 0 0 1 1 1 0 2 0 0 - 0 0 0 0 0 0 0 0 0 Euphausia superba 0 1 3 1 0 0 0 - 2 1 0 - - - 0 1 0 0 0 0 0 0 1 1 1 1 1 3 - 0 0 1 1 0 0 2 0 0 - 0 0 0 0 0 0 0 0 0 Anaspides tasmaniae 0 2 0 1 0 0 0 - 0 1 0 - - - 0 1 0 0 0 0 0 0 0 0 0 0 0 - - 0 0 0 1 0 0 2 0 0 - 0 0 0 0 0 0 0 0 0 Neognathophausia ingens 0 0 2 - 0 0 0 - 0 1 0 - - - 0 1 0 0 0 0 0 0 1 1 1 1 1 1 1 0 0 1 1 0 0 2 0 0 - 0 0 0 0 0 0 0 0 0 Heteromysis formosa 0 0 2 1 0 0 0 - 0 1 0 - - - 0 2 0 0 0 0 0 0 1 1 1 1 1 1 1 1 - 0 1 0 0 2 0 0 - 0 0 0 0 0 0 0 0 0 Stygiomysis holthuisi 0 0 3 1 0 0 0 - 0 1 0 - - - 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 - 0 1 0 0 0 0 0 - 0 0 0 0 0 - 0 0 0 Thetispelecaris remex 1 - - 1 0 3 0 - 0 1 0 - - - 2 2 0 0 0 0 0 0 3 3 0 0 0 0 - 0 1 0 0 0 0 2 0 0 - 0 0 1 1 0 0 0 0 0 Ingolfi ella tabularis 1 - - 1 0 2 1 2 0 1 0 - - - 2 2 0 0 0 0 0 1 - - - 1 - 0 0 1 0 2 2 0 - 0 0 2 0 0 - 0 0 2 Arrhis phyllonyx 1 - - 1 0 2 1 2 0 1 0 - - - 2 0 0 0 0 0 0 1 - - - 0 1 2 0 1 0 2 2 0 - 0 0 2 0 0 0 0 0 0 Protella gracilis 1 1 0 2 1 2 0 1 0 2 0 0 0 0 0 0 1 0 1 0 2 0 0 -Hyperietta stephenseni 1 - - 1 0 2 1 2 0 1 0 - - - 2 0 0 0 0 0 0 1 - - - 0 1 2 0 1 0 2 2 0 - 0 0 2 0 0 0 0 0 0 Gammarus troglophilus 1 - - 1 0 2 1 2 0 1 0 - - - 2 0 0 0 0 0 0 1 - - - 0 1 2 0 1 0 2 2 0 - 0 0 2 0 0 0 0 0 0 Tethysbaena argentarii 0 0 2 1 0 0 0 - A 1 0 - - - 2 2 0 0 0 0 0 0 2 2 2 2 2 2 2 0 1 0 0 0 0 2 0 0 - 0 1 0 1 0 - 0 0 2 Mictocaris halope 1 - - 1 0 0 0 - 0 1 0 - - - 0 2 0 0 0 0 0 0 2 2 2 2 2 - - 0 1 1 0 0 0 2 0 0 - 0 0 0 0 0 - 0 0 2 Spilocuma salomani 0 1 - - 0 1 0 - 0 1 0 - - - 1 - 0 0 0 0 0 0 - 0 0 0 2 - - 0 - 0 0 0 0 2 0 0 - 0 0 0 1 0 0 0 0 0 Diastylis sculpta 0 1 - - 0 1 0 - 0 1 0 - - - 1 - 0 0 0 0 0 0 - 0 0 0 2 - - 1 - 0 0 0 0 2 0 0 - 0 0 0 1 0 0 0 0 0 Spelaeogriphus lepidops 0 1 - 0 0 0 0 - 1 1 0 - - - 0 2 0 0 0 0 0 0 2 2 2 1 1 1 1 1 - 1 0 0 0 2 0 0 - 0 0 1 1 0 0 0 0 0 Apseudes bermudeus 0 1 - 0 0 3 0 - 0 1 0 - - - 3 0 0 0 0 0 0 0 3 3 - - - 1 - 0 0 0 0 2 0 0 - 0 1 1 1 0 0 0 0 0 Kalliapseudes sp. 0 1 - 0 0 3 0 - 0 1 0 - - - 3 0 0 0 0 0 0 0 3 3 0 0 0 0 - 1 - 0 0 0 0 2 0 0 - 0 1 1 1 0 0 0 0 0 Leptochelia sp. 0 1 - 1 1 0 0 - 0 1 0 - - - 3 0 0 0 0 0 0 2 - - - 1 - 0 0 0 0 0 0 0 - 0 1 1 1 0 0 0 0 3 Paratanais malignus 0 1 - 1 1 0 0 - 0 1 0 - - - 3 0 0 0 0 0 0 2 - - - 1 - 0 0 0 0 0 0 0 - 0 1 1 1 0 0 0 0 3 Tanais dulongii 0 1 - 1 1 0 0 - 0 1 0 - - - 3 0 0 0 0 0 0 2 - - - 1 - 0 0 0 0 0 0 0 - 0 1 1 1 0 0 0 0 3 Paramphisopus palustris 0 2 - 0 0 0 0 - 0 0 1 0 0 0 0 0 2 0 2 0 0 1 0 2 - - - 1 - 1 0 1 0 0 1 0 - 1 1 1 1 0 0 0 1 3 Colubotelson thomsoni 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 0 0 1 0 2 - - - 1 - 1 0 1 0 0 1 0 - 1 1 1 1 0 1 0 1 3 Stenasellus racovitzai 0 2 - 0 0 0 0 - 1 0 1 0 1 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 0 0 0 2 0 - 0 - 0 1 1 1 0 1 1 0 3 Caecidotea racovitzai 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 0 0 1 0 2 - - - 1 - 0 0 0 2 1 - 0 - 0 1 1 1 0 1 1 0 3 Asellus aquaticus 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 0 0 1 0 2 - - - 1 - 0 0 0 2 1 - 0 - 0 1 1 1 0 1 1 0 3 Stenetriid sp. 0 2 - 0 0 0 0 - 1 0 1 0 1 1 0 0 2 0 1 0 0 0 0 2 - - - 1 - 0 0 0 2 1 - 1 0 0 1 1 1 0 1 1 0 3 Janira maculosa 0 2 - 0 0 0 0 - 1 0 1 0 1 1 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 0 3 1 - 1 1 0 1 1 1 0 1 3 0 3 Joeropsis coralicola 0 2 - 0 0 0 0 - 1 0 1 0 1 1 0 0 0 2 0 1 1 0 0 2 - - - 1 - 0 0 0 3 1 - 1 1 0 1 1 1 0 1 3 0 3 Iathrippa trilobatus 0 2 - 0 0 0 0 - 1 0 1 0 1 1 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 0 3 1 - 1 1 0 1 1 1 0 1 3 0 3 Haploniscus nudifrons 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 1 0 2 0 1 1 0 0 2 - - - 1 - 0 0 0 4 1 - 0 - 0 1 1 1 0 1 3 0 3 Acanthaspidia drygalskii 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 0 3 1 - 1 1 0 1 1 1 1 1 3 0 3 Neojaera antarctica 0 2 - 0 0 0 0 - 1 0 1 0 0 1 0 0 0 2 0 1 1 0 0 2 - - - 1 - 0 0 0 3 1 - 1 0 0 1 1 1 0 1 3 0 3 Thylakogaster sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 2 3 1 - 0 - 0 1 1 1 1 1 3 0 3 Dendromunna sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 1 4 1 - 0 - 0 1 1 1 1 1 3 0 3 Eurycope sarsi 0 2 - 0 0 0 0 - 1 0 1 0 0 0 1 0 2 2 0 0 1 0 1 2 - - - 1 - 0 0 1 3 1 - 0 - 0 1 1 1 1 1 3 0 3 Ilyarachna antarctica 0 2 - 0 0 0 0 - 1 0 1 0 0 0 1 0 2 2 0 0 1 0 1 2 - - - 1 - 0 0 1 3 1 - 0 - 0 1 1 1 1 1 3 0 3 Munnopsis typica 0 2 - 0 0 0 0 - 1 0 1 0 0 0 1 0 2 2 0 0 1 0 1 2 - - - 1 - 0 0 1 3 - - 0 - 0 1 1 1 1 1 3 0 3 Betamorpha fusiformis 0 2 - 0 0 0 0 - 1 0 1 0 0 0 1 0 2 2 0 0 1 0 1 2 - - - 1 - 0 0 1 3 1 - 0 - 0 1 1 1 1 1 3 0 3 Mesosignum cf. usheri 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 0 2 0 ? 1 0 0 2 - - - 1 - 0 0 1 4 - - 1 1 0 1 1 1 0 1 3 0 3 Macrostylis sp.1 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 1 2 2 0 1 1 0 0 2 - - - 1 - 0 0 1 4 - - 1 0 0 1 1 1 0 1 3 0 3 Janirella sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 2 0 1 1 0 0 2 - - - 1 - 0 0 1 4 1 - 1 1 0 1 1 1 1 1 3 0 3 Ischnomesus sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 1 2 2 0 1 1 0 0 2 - - - 1 - 0 0 1 3 1 - 0 - 0 1 1 1 0 1 3 0 3 Eugerdella natator 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 1 2 2 0 1 1 0 2 2 - - - 1 - 0 0 1 3 1 - 1 0 0 1 1 1 0 1 3 0 3 Eugerda sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 1 2 2 0 1 1 0 2 2 - - - 1 - 0 0 1 3 1 - 1 0 0 1 1 1 0 1 3 0 3 Zonophryxus quinquedens 1 1 0 0 1 1 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 1 0 0 0 4 0 0 1 0 0 0 -Probopyrus pacifi ciensis 1 - - 1 0 0 1 1 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 2 0 0 - 0 1 0 0 0 0 0 0 1 Hemiarthrus abdominalis 1 - - 1 0 0 1 1 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 2 0 0 - 0 1 0 0 0 0 0 0 1 Paragnathia formica 1 - - 1 1 0 0 - 1 0 1 0 0 0 0 0 1 - - 0 0 0 - 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Riggia paranensis 1 - - 1 1 0 1 1 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Anilocra physodes 1 - - 1 1 0 1 1 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Cymodoce tattersalli 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 4 0 2 1 0 - 0 - 0 1 0 0 1 0 0 0 1 Campecopea hirsuta 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 4 0 2 3 1 - 0 - 0 1 0 0 1 0 0 0 1 Sphaeroma serratum 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 1 0 2 1 0 0 0 2 - - - 1 - 4 0 2 1 0 - 0 - 0 1 0 0 1 0 0 0 1 Cassidinidea sp. 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 4 0 2 3 0 - 0 - 0 1 0 0 1 0 0 0 1 Limnoria quadripunctata 0 2 - 0 0 0 0 - 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 1 0 0 0 0 1 0 - 0 1 0 0 0 0 0 0 1 Eurydice pulchra 1 - - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Excorallana quadricornis 1 - - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Natatolana albinota 1 - - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Aega antarctica 1 - - 1 1 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 1 0 0 0 0 0 0 - 0 1 0 0 0 0 0 0 1 Ligia oceanica 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Ligia italica 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Ligidium germanicum 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Haplophthalmus danicus 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Porcellio scaber 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Oniscus asellus 0 2 - 1 0 0 0 - 1 0 1 0 0 0 0 0 0 0 2 1 0 0 0 2 - - - 1 - 3 0 0 0 2 0 0 - 0 1 1 1 0 1 0 0 1 Paranthura nigropunctata 0 2 - 1 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 0 0 0 0 0 1 0 - 0 1 0 0 1 0 4 0 1 Cyathura carinata 0 2 - 1 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 0 0 0 0 2 - - - 1 - 0 0 0 0 0 1 0 - 0 1 0 0 1 0 4 0 1 Pygolabis humphreysi 0 2 - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 1 0 0 0 0 1 0 - 0 1 0 0 0 0 0 0 1 Idotea baltica 0 2 - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 2 0 1 2 2 - 0 - 0 1 0 0 1 0 0 0 1 Glyptonotus antarcticus 0 2 - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 2 0 1 2 2 - 0 - 0 1 0 0 1 0 0 0 1 Cleantis prismatica 0 2 - 0 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 2 0 1 1 0 - 0 - 0 1 0 0 1 0 0 0 1 Antarcturus spinacoronatus 0 2 - 1 0 0 1 0 1 0 1 0 0 0 0 0 2 0 2 1 0 0 0 2 - - - 1 - 2 0 1 2 0 - 0 3 0 1 0 0 1 0 0 0 1

Tab. 4. Continuation.

Character # 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 2 2 5 5 5 5 5 5 5 5 5 6 6 6 6 6 6 6 6 6 6 7 7 7 7 7 7 7 7 7 7 8 8 8 8 8 8 8 8 8 8 9 9 9 9 9 9 9 9 9 9 0 0 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 Gonodactylus viridis 0 0 1 0 1 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Penaeus semisulcatus 0 0 4 0 4 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Euphausia superba 0 0 1 0 1 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Anaspides tasmaniae 0 0 2 0 3 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Neognathophausia ingens 0 0 2 0 2 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Heteromysis formosa 0 0 2 0 2 - - - 1 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Stygiomysis holthuisi 0 0 3 0 3 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Thetispelecaris remex 0 0 4 0 4 - - - 1 - - - 0 0 0 0 0 0 1 0 0 0 2 0 Ingolfi ella tabularis 1 - - - - 2 5 0 5 - 0 - - - 1 - - - 0 2 0 0 0 0 0 1 0 0 1 1 Arrhis phyllonyx 1 - - - - 0 5 0 5 - 0 - - - 1 - - - 0 2 0 0 0 0 0 1 0 0 1 1 Protella gracilis 1 -Hyperietta stephenseni 1 - - - - 0 5 0 5 - 0 - - - 1 - - - 0 2 0 0 0 0 0 1 0 0 1 1 Gammarus troglophilus 1 - - - - 0 5 0 5 - 0 - - - 1 - - - 0 2 0 0 0 0 0 1 0 0 1 1 Tethysbaena argentarii 1 - - - - 2 - 1 - - 0 - - - 1 - - - 0 0 0 0 0 0 1 0 0 0 1 0 Mictocaris halope 0 - 3 0 3 - - - 1 - - - 0 0 0 0 0 0 1 0 0 0 0 0 Spilocuma salomani 0 0 1 0 1 - - - 1 - - - 0 0 0 0 0 0 1 0 0 0 1 0 Diastylis sculpta 0 0 1 0 1 - - - 1 - - - 0 0 0 0 0 0 1 0 0 0 1 0 Spelaeogriphus lepidops 0 0 1 0 1 - - - 1 - - - 0 0 0 0 0 0 1 2 0 0 1 2 Apseudes bermudeus 0 0 1 0 1 - - - 1 - - - 2 0 0 0 0 0 1 0 0 0 2 0 Kalliapseudes sp. 0 0 1 0 1 - - - 1 - - - 2 0 0 0 0 0 1 0 0 0 2 0 Leptochelia sp. 0 0 1 0 1 - - - 1 - - - 2 0 0 0 0 0 1 0 0 0 0 0 Paratanais malignus 0 0 1 0 1 - - - 1 - - - 2 0 0 0 0 0 1 0 0 0 0 0 Tanais dulongii 0 0 1 0 1 - - - 1 - - - 2 0 0 0 0 0 1 0 0 0 0 0 Paramphisopus palustris 1 - - - - 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 - 0 1 0 0 0 0 0 - 0 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 1 0 0 1 1 Colubotelson thomsoni 1 - - - - 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 - 0 0 0 0 0 0 0 - 0 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 1 0 0 1 1 Stenasellus racovitzai 1 - - - - 1 1 0 1 1 1 0 0 0 0 0 1 0 1 1 0 1 0 3 1 0 0 1 0 1 0 1 0 0 0 0 0 0 0 1 2 0 0 0 0 0 0 1 0 0 1 1 Caecidotea racovitzai 1 - - - - 1 1 0 1 1 1 0 0 0 0 0 1 0 1 1 0 1 1 2 0 0 2 2 0 1 0 1 0 0 0 0 0 0 0 3 2 0 0 0 0 0 0 0 0 0 1 1 Asellus aquaticus 1 - - - - 1 1 0 1 1 1 0 0 0 0 0 1 0 1 1 0 1 1 2 0 0 2 2 0 1 0 1 0 0 0 0 0 0 0 3 2 0 0 0 0 0 0 0 0 0 1 0 Stenetriid sp. 1 - - - - 1 1 1 1 1 1 0 0 0 1 0 2 0 1 1 0 1 0 4 0 0 1 1 1 2 1 1 0 0 1 0 0 0 0 1 2 0 0 0 0 0 0 0 0 0 1 0 Janira maculosa 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 2 0 0 1 2 2 0 0 0 0 0 0 0 0 0 1 0 Joeropsis coralicola 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 2 0 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Iathrippa trilobatus 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 0 0 0 1 2 2 0 0 0 0 0 0 0 0 0 1 0 Haploniscus nudifrons 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 2 1 1 2 2 2 0 - 0 1 - - 0 0 1 0 Acanthaspidia drygalskii 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 1 1 1 1 1 3 2 2 0 - - 1 - - 0 0 1 0 Neojaera antarctica 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 1 0 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Thylakogaster sp. 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 1 0 1 1 2 2 2 0 - - 1 - - 0 0 1 0 Dendromunna sp. 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 2 2 0 1 1 2 3 2 0 - - 1 - - 0 0 1 0 Eurycope sarsi 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 1 2 0 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Ilyarachna antarctica 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 1 1 1 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Munnopsis typica 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 0 2 0 1 1 2 2 2 0 - - 1 - - 0 0 1 0 Betamorpha fusiformis 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 0 0 1 2 0 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Mesosignum cf. usheri 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 2 1 1 2 2 2 0 - - 1 - - 0 0 1 0 Macrostylis sp.1 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 2 1 1 2 2 2 0 - - 1 - - 0 0 1 0 Janirella sp. 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 1 2 1 0 1 2 2 2 0 - - 1 - - 0 0 1 0 Ischnomesus sp. 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 1 1 1 2 2 2 0 0 0 A 0 0 0 0 1 0 Eugerdella natator 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 2 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Eugerda sp. 1 - - - - 1 1 1 1 1 1 1 1 1 1 1 2 1 1 2 0 1 0 5 1 1 1 1 1 3 1 1 1 0 2 2 2 1 1 2 2 2 0 0 0 0 0 0 0 0 1 0 Zonophryxus quinquedens 0 1 1 0 1 -Probopyrus pacifi ciensis 0 0 1 0 1 0 1 2 1 0 0 0 0 2 0 0 1 -Hemiarthrus abdominalis 0 0 1 0 1 0 1 2 1 0 0 0 0 2 0 0 1 -Paragnathia formica 0 0 1 0 1 - - - 0 1 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Riggia paranensis 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Anilocra physodes 0 0 1 0 1 - - - 0 0 0 - 0 0 ? - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Cymodoce tattersalli 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 1 0 0 0 0 0 2 0 1 1 2 Campecopea hirsuta 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 1 0 0 0 0 0 2 0 1 1 2 Sphaeroma serratum 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 1 0 0 0 0 0 2 0 1 1 2 Cassidinidea sp. 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 1 0 0 0 0 0 2 0 1 1 2 Limnoria quadripunctata 0 0 1 0 1 - - - 0 0 0 - 0 0 0 0 0 0 - - - 0 0 0 0 0 0 0 1 0 0 1 2 Eurydice pulchra 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Excorallana quadricornis 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Natatolana albinota 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Aega antarctica 0 0 1 0 1 - - - 0 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 0 0 0 2 0 0 1 2 Ligia oceanica 1 - - - - 1 1 0 1 1 0 0 0 - 0 - 0 0 0 0 0 0 - 0 0 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 1 0 0 0 1 0 0 1 1 Ligia italica 1 - - - - 1 1 0 1 1 0 0 0 - 0 - 0 0 0 0 0 0 - 0 0 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 1 0 0 0 1 0 0 1 1 Ligidium germanicum 1 - - - - 1 1 0 1 1 0 0 0 - 0 - 0 0 0 0 0 0 - 0 0 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 0 1 0 0 1 0 0 1 1 Haplophthalmus danicus 1 - - - - 1 1 0 1 0 0 0 0 - 0 - 0 0 0 0 0 0 - 0 1 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 0 2 0 0 1 0 0 1 1 Porcellio scaber 1 - - - - 1 1 0 1 0 0 0 0 - 0 - 0 0 0 0 0 0 - 0 1 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 0 2 0 0 1 0 0 1 0 Oniscus asellus 1 - - - - 1 1 0 1 0 0 0 0 - 0 - 0 0 0 0 0 0 - 0 1 0 0 0 0 - - - 1 0 0 0 0 1 0 3 2 0 0 0 2 0 0 1 0 0 1 0 Paranthura nigropunctata 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 1 0 0 2 0 0 1 2 Cyathura carinata 0 0 1 0 1 - - - 0 0 0 - 0 0 0 - 0 0 - - - 0 0 0 0 1 0 0 2 0 0 1 2 Pygolabis humphreysi 0 0 1 0 1 - - - 0 0 0 - 1 0 0 - 0 0 - - - 2 0 - 0 0 0 0 0 0 0 1 1 Idotea baltica 0 0 1 0 1 - - - 0 0 0 - 0 0 0 0 0 0 - - - 0 1 1 0 - 1 - - 0 0 1 2 Glyptonotus antarcticus 0 0 1 0 1 - - - 0 0 0 - 0 0 0 0 0 0 - - - 0 1 1 0 - 1 - - 0 0 1 2 Cleantis prismatica 0 0 1 0 1 - - - 0 0 0 - 0 0 0 0 0 0 - - - 0 1 1 0 0 0 0 2 0 0 1 2 Antarcturus spinacoronatus 0 0 1 0 1 - - - 0 0 0 - 0 0 0 0 0 0 - - - 0 1 1 0 0 0 0 2 0 0 1 2

the parameters: the families Bopyridae and Cymot-hoidae both had the highest monophyly frequency (frequency of topological congruence) of 93% and 100%, respectively. Other taxa, such as the Phreato-icidea and the Asellidae had a monophyly frequency of 71%. The large superfamily Janiroidea (18 taxa) and the large deep-sea family Munnopsidae showed a relatively high frequency of 50% of the parameter sets. This is not a general result because some family-level taxa (e.g., the Ligiidae) were found in none of the analyses. The absence of a Sphaeromatidae clade in all analyses was caused by the sequence from one species (Cymodoce tattersalli), a result that should be reconfi rmed with new sequence data. The relation-ships of the families Cirolanidae and Aegidae may need to be re-assessed because a species of Aegidae (Aega antarctica) was frequently nested within a clade of two cirolanid species (Eurydice pulchra and Natatolana meridionalis).

Im Dokument The Phylogenetic Position of the Isopoda in the Peracarida (Crustacea: Malacostraca) (Seite 21-26)