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Brändli, U. B. (2005). Tree species diversity. In Swiss Agency for the Environment,Forest and Landscape SAEFL,Berne & Swiss Federal Institute WSL,Birmensdorf (Eds.), Forest Report 2005. Facts and Figures about the Condition of Swiss Forests (pp. 74-75).

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74 FOREST REPORT 2005

4.1a T ree species diversity

Switzerland’ s surface would naturally be covered mostly by beech and spruce forests, which contain relatively few species.

Most Swiss forests today contain mainly commercially interesting species, such as spruce, fir , beech and ash.

The species diversity of trees in Swiss forests has increased during the past ten years by 4%.

This also promotes the diversity of other plants and animals.

Promoting competitively weaker, but ecologically more valuable tree species increases biodiversity.

Number of species

The twoSwissNationalForest In- ventories to date have revealed a pleasing trend: between 1985 and 1995 the species diversity of trees increased by 4%.The in- crease was largest south of the Alps (10%) and smallest in the Central Plateau (1%). The total area of monotonous forest also decreased slightly during the same period.This isanother pos- itivedevelopment becausemon- otonous forestscontain far few- eranimalandplant species than mixed stands. Programmes pro- moting tree diversity therefore make ecologicalsense.Thebeech andsubalpine spruce forests that today dominate the Swiss land- scape naturally contain rather few species. Forests rich in dif- ferent tree species, on the oth- er hand, which are often creat- ed through human intervention, provideamultifacetedmosaicof habitats. Silvicultural treatments can today promotecompetitive- ly weak tree species that require light in the sameway as the tra- ditionalforest management prac- tices,coppice forestand coppice with standards,used todo.Thus Zurich’s Sihl Forest, for exam- ple, which was once intensively used,is today richer in tree spe- cies thana beech virgin forest in theCarpathians.The situation is, however, different for those in- sects and fungi that depend on dead wood: they find betterliv- ingconditions invirginandnat- ural forests (> 4.5 Dead wood).

Nevertheless, around 1300 spe- cies of plants, including 12 in- digenousconifersand 43 indig- enousbroadleaf trees,still grow in Swiss forests. However, 70%

ofSwiss forest ismadeup ofjust three species of tree:spruce(40%

of the trees counted in 1995), beech(18%)andfir(11%).

The individual tree spe- cies have developed different- ly.The number of spruce,beech, fir, Scotch pine (Pinus sylves- FURTHERINFORMATION

Swiss Federal Research Institute WSL 8903 Birmensdorf

National Forest Inventory Research Dept. LANDSCAPE Section Landscape inventories (0) 44/ 739 23 43

Swiss Agency for the Environment, Forests and Landscape SAEFL 3003 Bern

Forest Agency

Section Forest utilization and timber industry

Section Forest conservation and biodiversity

(0) 31/ 324 77 78

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75 4BIOLOGICAL DIVERSITY

tris), common oak and sessile oak went downconsiderably be- tween1985and1995,while those of sycamore,linden, rowan and whitebeam went up. The abun- dance of wych elm fell by 30%

when an introduced fungal dis- ease spread.In contrast, the oc- currence of English yew, a rare species,has increasedslightly.

SAEFL started a project in 1997 togetherwith the ETH Zu- rich to promote relatively rare tree species(SEBA),suchas yew, wildservice tree(Sorbus tormina- lis)or small-leaved linden (Tilia cordata).The aim of the project is to increase the populations of these species and to raise forest owners’ and managers’ aware-

Forest management

Aforest rich in tree species is of- ten the result ofcareful manage- ment. If no trees are felled, the forest will become increasingly older,andshade-tolerant species suchasbeech,firandsprucewill eventually dominate at the ex- pense of species that needlight.

Left to its own devices,a forest willgrowdenser overalongperi- od and the treelayerwillbecome more uniform. Felling trees, on the other hand,bringslight into the forest and thus promotesdi- versity. Although at most sites natural regeneration is prefera- ble,planting young treescan en- hancebiodiversity solongas the treesarelocalspecies.Nowadays, many forestsare less intensively managed,so that fewer regener- ationareasarebeingestablished and fewer stands thinned. This has, in the short and medium term,anegative effect on biodi- versity.

4.1a.1

Mean number of tree species

Trees12 cmin diameter andlarger on the500m2 sample sites of theNationalForest Inventory NFI2.

Switzerland: 25

■ up to 2.0

■ 2.1–2.3

■ 2.4–2.6

2.7–2.9

≥ 3.0 3.0

3.0 3.0

2.8

2.8

2.8 2.5

2.5

2.5 2.4 2.3

2.3 2.2

1.8

4.1a.2

Distribution of tree species

Trees12 cmin diameter andlarger on the500m2 sample sites of theNationalForest Inventory NFI2.

≥ 3 species

3 species

2 species

1 species

LFI1 LFI2 LFI1 LFI2 LFI1 LFI2 Broadleaf forest area Conifer forest area Total 100

75 50 25 0

[%]

ness. In addition to the SEBA species, the government is pro- moting the cultivation of oaks, especially on areas affected by the stormLothar.

The diversity of tree species in the Central Plateau is, with- out human intervention, great- er than it is in themountains,as biodiversity decreases with alti- tude.Speciesdistributedinareas with so-called Atlantic and sub- Mediterranean climates thrive betterat loweraltitudes because of the longer vegetation period and milder winter. Species be- longing to thiscategory are oaks, Norway,Italianandfieldmaples, elmsandwildservice trees.

The diversity of tree species in the Central

Plateau is, without human intervention,

greater than it is in the mountains, as bio-

diversity decreases with altitude.

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